Accepted Name - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

ISSN: 2386-9666 - Last Updated: 11 August 2026

Accepted Name:

Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298[-300], t. 21 (1807) (Desfontaines 1807: 298-300, t. 21; 1808, 16, t. 10; Sprengel 1825: 818; Don 1838: 633; Piwowarczyk & al. 2019: 18)

Synonyms:

Homotypic synonyms:

≡ Phelypaea Orientalis, flore coccineo Tournef., Corollarium: 47 t. 479 (1703) [pre-Linnaean name, not validly published] (Desfontaines 1807: 298, "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Desfontaines 1808: 16, "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Don 1838: 633; Stapf 1915: 285; Cullen 2010: 380)

≡ Lathraea phelypaea var. β [unnamed] L., Sp. Pl. 2: 606 (1753 [1 May 1753]) ["β Orobanche orientalis, flore coccineo. Tournef. cor. 47."], nom. illeg. (Stapf 1915: 286)

≡ Anoplon tournefortii (Desf.) G. Don, Gen. Hist. 4: 633 (1837-1838) (Piwowarczyk & al. 2019: 18)

≡ Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) ["A. Tournefortii Walp. rep. bot. 3 p. 48, ex parte"]; [Walp., Repert. Bot. Syst. 3: 481 (1844 [12-14 Sep 1844]) (Beck 1930: 44; Piwowarczyk & al. 2019: 18)]. Obs. "basi vaginis elongatis inaequalibus involuti. Calyx ... villosus ... (v. s. in h. Bentham.)" (Reuter 1847: 43)

≡ Aeginetia tournefortii (Desf.) [Juss. sec. Steudel] Beck in Engl., Pflanzenr. 96: 44 (1930), pro syn.

≡ Diphelypaea tournefortii (Desf.) Nicolson in Taxon, 24(5-6): 654 (1975 [Nov 1975]), nom. illeg. (Piwowarczyk & al. 2019: 18)

inicio / top of page

Heterotypic synonyms:

? = Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name, not validly published]. Syn.: ≡ Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]) (Linnaeus 1753: 606; Don 1838: 635); = Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811) (Endlicher 1839: 727)

? = Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]). Syn.: ≡ Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name] (Linnaeus 1753: 606; Don 1838: 635); ≡ Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838); = Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811). Ind. loc.: "Habitat in Oriente" (Linnaeus 1753: 606); "Asia minor" (Sprengel 1825: 818); "H. Native of the Levant" (Don 1838: 635; Lathraea anblatum in the Linnaean Typification Project, accessed Mar 2012). Obs. According to Endlicher (1839: 727), Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) is conspecific with Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811). According to Reuter (1847: 43), Anblatum tournefortii G. Don is conspecific with Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807). According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753) [this synonym seems correct on the base of Don's description]

? = Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838) (Reuter 1847: 43). Obs. According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753)

? = Aeginetia coccinea Juss. ex Reut. in DC., Prodr. 11: 44 (1847), pro syn. Syn.: = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 44 (1847) (Reuter 1847: 44)

inicio / top of page

= Anoplanthus coccineus var. peduncularis Boiss., Fl. Orient. 4(2): 494 (1879 [Apr-May 1879]) ["A. coccineus β peduncularis"] (Piwowarczyk & al. 2019: 18). Syn.: = Phelypaea Orientalis, flore coccineo Tournef., Corollarium: 47 t. 479 (1703); = Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298, t. 21 (1807); = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) (Boissier 1879: 494; Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]). Ind. loc.: Turkey: "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia] (Tourn!, Huet!, Calv!), in Kurdistaniâ (Barré de Lancy [vice-consul de France à Mersine et Adana]! [see P. boissieri], Armeniâ Rossicá (Buhse!), in pratis montosis Georgiae Caucasicae (Hoh!)". Ic.: "Phelypaea Tournefortii Desf. Cor. tab. 10" (Boissier 1879: 494). Obs. "Squamae ad basin caulis superne longe nudis magis congestae" (Boissier 1879: 494)

inicio / top of page

Ind. loc.: Turkey: "Il est originaire d'Arménie [Turkish Armenia / Armeniâ Turcicâ / Western Armenia]" (Desfontaines 1807: 300); "ad radices Pyrethri myriophylli C.A. Meyer [= Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. [= Tanacetum sp. ?] ... Armenien: Provinz Erzerum [Erzurum, Anatolia] (Tournefort..." (Beck 1930: 44; Novopokrovsky & Tzvelev 1958[2000]: 30[28]; Nicolson 1975: 652 "variety B [β]"; Cullen 2010: 380).

Typus / Type: "Type:—Turkey (lectotype designated here):—Phelypaea Orientalis, flore coccineo [Arménie, nowadays Erzurum Province, Turkey], Tournefort (1703: t. 479!). Turkey (epitype designated here):—Armenia, im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?], inter Baiburt [Bayburt] et Erzeroum [Erzurum], Maio 1853, Huet de Pavillon [Plantae Orientales exsiccatae] (P-02970941!)." (Piwowarczyk & al. 2019: 18, sub D. tournefortii).

N.B. "that no specimens of the Phelipaea Tournefortii now remain in his Herbarium, or are known to exist in any other collection" (Lambert 1811: 261); "No specimen of this species could be found in the Tournefort herbarium (P)" (Gilli 1982: 2); "There is, apparently, no specimen in the Tournefort herbarium, but Aubriet, the artist who accompanied Tournefort, made an illustration which appears as t. 479 of the Corollarium, and can be regarded as the type material; it is clearly the plant recognised here as Phelypaea tournefortii." (Cullen 2010: 380; Piwowarczyk & al. 2019: 18, sub D. tournefortii).

Designated by: Piwowarczyk & al. in Phytotaxa 386(1): 18 (2019).

Host: Apparently a parasite exclusively of Tanacetum species (Annotated Checklist of Host Plants of Orobanchaceae, accessed Aug 2009).

Asteraceae Bercht. & J. Presl (Compositae Giseke)

Tanacetum L. (Reuter 1847: 43, sub Anoplanthus tournefortii (Desf.) Reut.; Fischer 1852: 107, sub Anoplanthus Tournefortii; Beck 1930: 44, sub P. tournefortii; Novopokrovsky & Tzvelev 1958[2000]: 30[28], sub P. tournefortii; Gilli 1982: 2, sub P. tournefortii; Piwowarczyk & al. 2019: 19, sub D. tournefortii).

Tanacetum argyrophyllum (K. Koch) Tzvelev [≡ Tanacetum polycephalum subsp. argyrophyllum (K. Koch) Podlech ; = Tanacetum myriophyllum Willd., nom. illeg.; = Achillea bipinnata L.; – Pyrethrum myriophyllum auct. plur.]. Armenia (Piwowarczyk & al. 2019: 19, sub D. tournefortii).

Tanacetum aureum (Lam.) Greuter, M. V. Agab. & Wagenitz [≡ Achillea aurea Juss. ex Lam.; = Tanacetum chiliophyllum (Fisch. & C.A. Mey.) Sch. Bip.; = Pyrethrum chiliophyllum DC.]. Armenia (Piwowarczyk & al. 2019: 19, sub D. tournefortii), Turkey (E 01283801 cf.; K 004381263 cf.).

Tanacetum canescens DC. [≡ Pyrethrum canescens (DC.) Boiss.]. Armenia (Piwowarczyk & al. 2019: 19, sub D. tournefortii).

Tanacetum polycephalum Sch. Bip. (s.l.). Armenia (Piwowarczyk & al. 2019: 19, sub D. tournefortii), Azerbaijan, Nakhchivan (Bushe ex Fischer 1852: 107, sub Anoplanthus Tournefortii; Bushe ex Stapf 1915: 293, sub P. tournefortii; Beck 1930: 44, sub P. tournefortii; Novopokrovsky & Tzvelev 1958[2000]: 30[28]), Turkey (K 004381267).

Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech [≡ Pyrethrum duderanum Boiss.; = Pyrethrum myriophyllum sensu C.A. Mey. [non Tanacetum myriophyllum Willd., nom. illeg.]. Azerbaijan, Nakhchivan (Bushe ex Fischer 1852: 107, sub Anoplanthus Tournefortii; Bushe ex Stapf 1915: 293, sub P. tournefortii; Beck 1930: 44, sub P. tournefortii; Novopokrovsky & Tzvelev 1958[2000]: 30[28], sub P. tournefortii), Turkey (J. Brant in K 004381267).

Comments:

Morphological observations: "P. Scapo basi involuto, villoso ... Calice ... velues ... Le Phelipaea de Tournefort a les tiges nues et les divisiones de la corolle sont circulaires et non ovales" (Desfontaines 1807: 298-300). Key to these species (Stapf 1915: 290; Novopokrovskii & Tzvelev 1958[2000]: 28[26]; Cullen 2010: 383-384); pollen morphology (Zare & al. 2014); key (Piwowarczyk & al. 2019: 19).

Phylogenetic observations: Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57 [x = 19, sub Phelypaea (syn. Diphelypaea)], 61 f. 4 ["Phylogenetic relationships", sub Phelypaea], 63 (2005); "Recent molecular phylogenetic studies suggest two major lineages, informally called the Orobanche-group and the Phelipanche-group (Schneeweiss et al., 2004a). The first includes Orobanche sect. Orobanche and the small SW Asian [and SE Europe] genus Phelypaea (listed as Diphelypaea in Schneeweiss et al., 2004a), while the second includes the remaining sections of Orobanche (sects. Gymnocaulis, Myzorrhiza, and Trionychon). The circumscription of these groups is well supported by the distribution of chromosome base numbers, these being x = 19 in the Orobanche-group and x = 12 in the Phelipanche-group (Schneeweiss et al. 2004b)" (Weiss-Schneeweiss & al., 2006; Schneider, 2016: f. 1).

Taxonomic observations:

Phelypaea L., Op. Var.: 237 (1758) is the correct name for the genus, because the previous names were not validly published according to ICN (Turland & al. 2018).

Phelypaea Hill in Scott, Suppl. Chambers Cyclop. (1753) [not validly published] (Lonchamp in INRA-MNHN 1999; HUH id=16687, accessed Jan 2020; Turland & al. 2018: Art. 34, "suppressed work" (see App. I)). Type Name: Lathraea phelypaea L., Sp. Pl.: 606 (1753)

Phelypaea Hill in Owen, Dict. Arts Sci. 3: 2416 (1755 [1 Mar 1755]) [not validly published] ["Phelypaea, in botany, a genus of the didynamia-angiospermia..."] (Reveal 1991: 228, "The Hill genus name is now the oldest one available"; Turland 2018: Art. 34, "suppressed work" (see App. I)). NT [Neotype]: Lathraea phelypaea L., Sp. Pl.: 606 (1753) (Reveal 1991: 228); ≡ Cistanche phelypaea (L.) Cout. (1913)

Phelypaea P. Browne, Civ. Hist. Jamaica: 269 (1756 [Mar 1756]) [not validly published] (Shenzhen Code - Art. 38.6 Ex. 13, "In publishing "Phaelypea" without a generic description or diagnosis, Browne (Civ. Nat. Hist. Jamaica: 269. 1756) included and described a single species [Phelypaea erecta P. Browne], but he gave the species a phrase name (McNeill & Greuter 2016) not a validly published binomial. Art. 38.5 does not therefore apply and "Phaelypea" is not a validly published name.") (Turland & al. 2018: Art. 38.6 Ex. 13).

Phelypaea L., Op. Var.: 237 (1758), non Phelypaea Hill (1755) nec Phelypaea P. Browne (1756) ["Phaelypae"] (Reveal 1991: 228; Nicolson 1975: 653). Type Name: Phelypaea tournefortii Desf. (1807); ≡ Diphelypaea tournefortii (Desf.) Nicolson (1975). Obs. Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57, 58, 61 f. 4, 63 (2005); "vide Dandy, Ind. Gen. Vasc. Pl. 1753-74 (Regn. Veg. li.) 7 (1967)". Replaced synonym of: Diphelypaea Nicolson Taxon 24(5-6): 654. 1975 (view IPNI entry, accessed Feb 2008). "Dr R.K. Brummitt of the Royal Botanic Gardens, Kew tells me that he regards the name Phelypaea as the correct one for the genus." (Cullen 2010: 380).

Diphelypaea Nicolson in Taxon, 24(5-6): 654 (1975), nom. nov. [nom. illeg.]. Syn.: ≡ Phelypaea L., Op. Var.: 237 (1758). Type Name: Diphelypaea tournefortii (Desf.) Nicolson (1975) [Phelypaea tournefortii Desf. (1807)]. IPNI: "without full basionym ref.". The reference to the place of publication of the basionym in "References"! (Nicolson 1975).

Chromosomatic number: n = 19, 2n = 38 (Schneeweiss & al. 2004, sub Diphelypaea; Carlón & al. 2005: 57, sub Phelypaea [between Bayburt and Erzurum, NE Anatolia, Turkey]).

Ic.: a, Tournefort, Corollarium: 47 t. 479 (1703), sub P. orientalis, flore coccinea; b, Desfontaines 1807: t. 21 et 1808: 16, t. 10, sub Phelypaea tournefortii; c, Beck in Engl., Pflanzenr. 96: 43 f. 9 (1930, sub P. Tournefortii); d, habit (Muradiye, Van, Turkey [May 2009, photo by P. Coulot]); e, flower (ibidem); f, calyx (ib. ["Calyx ... villosus" (Reuter 1847: 43)]); g, scale-leaves (ib. ["Squamae ad basin caulis superne longe nudis magis congestae" (Boissier 1879: 494)]); h, habit (P. tournefortii eastern Anatolia, Jul 2002 by G. Schneeweiss); i, flower (P. tournefortii eastern Turkey, by L.J. Musselman); j, sheet (Turkey in Demirkuş 2007); k, Novopokrovskij & Tzvelev (1958[2000]: 30[37] t, 3 f. 5); l, D. tournefortii vs. D. coccinea vs. D. boissieri (P, habits and calyces); n, Piwowarczyk & al. 2019: 18 f. 9 A-F, sub D. tournefortii.

Distribution: Eurasian distribution (Armenia, Azerbaijan, Georgia [needs be confirmed], Iran, Turkey).

Eurasian distribution (Domina & Raab-Straube 2010, sub Phelypaea tournefortii, accessed Jul 2010).

inicio / top of page

Specimens examined and literature records.

Armenia / Hayastan / Armenien / Arménie / Армения (Stapf 1915: 293, sub Anoplanthus coccineus var. nigrovittatus; Beck 1930: 44, sub P. tournefortii; Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii)

Aragatsotn Province / Арагацотнская область (Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii): 1. "... / 11^th Iter Mediterraneum of OPTIMA / Armenia, June 2002 / ... / Phelураea tournеfortii Desf. / Armenia: Kotayk province, Nairi distr., c. 3 km N Yeghvard, mountain Arailer [Ara / Араилер] / "Tsahhkevank", subalpine shrubland and meadows with rocky outcrops / 40°22'60.0"N 44°16'60.0"E 40.383330, 44.283330 [near Antarut / Антарут and Byurakan / Бюракан (Ashtarak Municipality)] / 29-06-2002 / leg. Fayvush et al. / det. Fayvush (ERE) 2006 / Nº Rec. It. 2373". – 2. "B 10 0469878 / ! / confirm.! Hulich & & S. Rätzel / 9.02.2022" (https://jacq.org/detail/490922 - http://herbarium.bgbm.org/object/B100469878 - B 10 0469878 - https://www.gbif.org/occurrence/893482736;  rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base, stem and calyx with conspicuous hairs); "Арагацотн [Aragatsotn, 40°21'25.1"N 44°23'41.5"E 40.356960, 44.394860, Kasagh canyon, near Ohanavan / Оганаван (Ashtarak Municipality)], 14 May 2023" (https://www.inaturalist.org/observations/161679571 - https://www.gbif.org/occurrence/4116313080 by https://www.inaturalist.org/people/aleksandr_volkov [Aleksandr Volkov], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, stem and calyx clearly hairy, apparently on Tanacetum); "9747+GP, Антарут 0213 [40°21'18.3"N 44°15'49.1"E 40.355080, 44.263650, Antarut / Антарут (Ashtarak Municipality)], 24 May 2025" (https://www.inaturalist.org/observations/285972371 by https://www.inaturalist.org/people/aleksandr_volkov [Aleksandr Volkov], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, stem and calyx clearly hairy, apparently on Tanacetum); "Aragatsotn [40°21'53.0"N 44°15'12.5"E 40.364720, 44.253480, near Antarut / Антарут (Ashtarak Municipality)], 27 May 2026" (https://www.inaturalist.org/observations/372428002 by https://www.inaturalist.org/people/ibidorhynchastruthersii, sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, scales only at the base, stem and calyx clearly hairy, apparently on Tanacetum); "Aragatsotn [40°23'16.4"N 44°23'56.9"E 40.387880, 44.399130, near Saghmosavan / Сагмосаван (Ashtarak Municipality)], 7 Jun 2026" (https://www.inaturalist.org/observations/369642602 by https://www.inaturalist.org/people/alvaro_antnio [Álvaro António], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, stem conspicuously hairy, apparently on Tanacetum).

Ararat Province / Араратская область (Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii): "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / В 2 км к югу от с. Кярки [2 km to the south of the village of Karki / Kərki / Kyarki / Tigranashen / Тигранашен (Садаракский район / Sadarak District)] можжевеловое редколесье, 1400 м. / 29 мая 1973 г. [29.05.1973] Н. Шведчикова [N. Shvedchikova]" (https://plant.depo.msu.ru/open/public/item/MW0723306 - MW 0723306.jpg - https://www.gbif.org/occurrence/1697840797; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.); "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / Окрестности Советаmена [Surroundings of the Council (Vedi)]. / Н. Шведчикова [N. Shvedchikova] 30 мая 1973 [30.05.1973] г. 7782" (https://plant.depo.msu.ru/open/public/item/MW0723307 - MW 0723307.jpg - https://www.gbif.org/occurrence/1697840788; rev. Ó. Sánchez Pedraja Apr 2018); "Plants of Armenia / ... / Flile NY: 288.0 / Phelipaea coccinea (M. Bieb.) Poir. / det. K. Tamanyan, / Armenia. 3 km S from Lusashoh village [Lusashogh / Garaxach / Лусашох], 39º49'72N, 44º59'195E / 1760 m. (... / ...). Mountain steppe. / Kamilla Tamanyan 10-2005  28 May 2005 / George Fayvush / The New York Botanical Garden (NY) / in collaboration with / Institute of Botany, Armenian Academy of Sciences (ERE)" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p03931901 - P 03931901.jpg - https://www.gbif.org/occurrence/607811327; rev. Ó. Sánchez Pedraja Apr 2018, sub  P. tournefortii); idem (http://specimens.kew.org/herbarium/K004381232 - K 004381232.jpg - https://records.data.kew.org/occurrences/7b20039c-0610-4b31-a98a-84961a1d8078 - https://www.gbif.org/occurrence/5060755061; rev. Ó. Sánchez Pedraja Aug 2026, sub  P. tournefortii); "Цветущее растение. Армения, Гарни, Хосровский заповедник [Garni, Khosrov Reserva] 18.05.2010" (http://www.plantarium.ru/page/image/id/107519.html by Ваэ Мартиросян [Vahe Martirosyan], 2011 in Plantarium [Diphelypaea tournefortii, on Tanacetum sp.] and iNaturalist; accessed Feb 2013); "Khosrov reserve 2295 m a.s.l., 5 June 2007" (https://www.inaturalist.org/observations/585400 - http://www.inaturalist.org/photos/744571 by Roberto Sindaco, sub Diphelypaea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jan 2015, the photo seems to correspond to P. tournefortii on Tanacetum sp.); "2WP8+PPV Natuurreservaat Khosrov [Khosrov Forest State Reserve, 40°07'23.5"N 44°46'07.0"E 40.123190, 44.768600, Mets Gilanlar], 19 May 2026" (https://www.inaturalist.org/observations/363420245 - https://www.gbif.org/occurrence/6353290941 by https://www.inaturalist.org/people/stijnvanacker, sub Phelypaea coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii on Tanacetum sp.); "Urtsalanj [39°48'57.0"N 44°59'32.2"E 39.815830, 44.992270, near Zovashen / Keshishveran (Ararat Municipality)], 1 Jun 2026" (https://www.inaturalist.org/observations/367521153 - https://www.gbif.org/es/occurrence/6352420450 by https://www.inaturalist.org/people/axelgosseries [Axel Gosseries], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base, stem and calyx with conspicuous hairs).

Gegharkunik Province / Гехаркуникская область (Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii): "Herbarium Universitatis Mosquеnsis (MW) / P. Smirnow. Pl. Armeniae ross. a. 1929. / Anoplanthus coccineus Walp. / Окрестн. оз. Гокча [lake Gokcha / Lake Sevan / Севан] / Дарачичаг [Darachig / Drakhtik / Драхтик (Shoghakat Municipality / Шохакат)] / Щебнистый склон 2200 м / 19.VI.1929. / № 843 / Собр. П. Смирнов [P. Smirnov]" (https://plant.depo.msu.ru/open/public/item/MW0723296 - MW 0723296.jpg - https://www.gbif.org/occurrence/1697840731; rev. Ó. Sánchez Pedraja Apr 2018, sub  P. tournefortii); "im Distr. Novobajazet [Novo Bajazet / Новобаязет / Nor Bayezet / Novyi Bayazet, nowadays Gavar / Kyavar / Гавар (Gavar Municipality)] um Solak, Berezovaja butka, Konstantinowska (E. Mylow!), bei Station Eilar [Dzak] (A. Fomin!)" (Beck 1930: 44, sub P. tournefortii); "Diphelypaea tournefortii, filmed in the Tarachichaki [Tsaghkunk / Varazhnunik] (town) ..." (http://ic.pics.livejournal.com/vahemart/27418323/85148/85148_original.jpg by Ваэ Мартиросян [Vahe Martirosyan], 2011 in vahemart.livejournal.com; accessed Jan 2015).

Kotayk / Kotaik Province / Котайкская область (Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii): "Caucasus, Gouv. Eriwan, St. Eyliar [Eilar / Eylar / Ellar / Элар, currently part of Abovyan / Абовян (10 km northeast of the city of Yerevan / Erywan / Jerewan / Ереван)], on Pyrethrum myriophyllum, Thomin (in Acta Hort. Tifl. vi. II. (1902) 56)" (Stapf 1915: 293, sub Anoplanthus coccineus var. nigrovittatus) [Pyrethrum myriophyllum is the typical host of D. tournefortii]; "Distr. Eriwan [Erywan / Yerevan / Jerewan / Ереван]: bei Hilar [Eilar / Eylar / Ellar / Элар, currently part of Abovyan / Абовян], zwischen Garyna und Arabat (Bayern)" (Beck 1930: 44, sub P. tournefortii); "Herbarium Universitatis Mosquеnsis / Plantae Caucasicae Collaboratoribus Horti / Botanici Universitatis Mosquеnsis Lectaе / Phelураea tournеfortii Desf. / Армения, Разданский р-н [Hrazdan raion], / хр. Цахкунец [Tsaghkunyats Mountains], басс. р. Мармарик [Marmarik], / окр. пос. Цахкадзор [Tsaghkadzor / Tsakhkadzor] / М.Г. Пименов Е.В. Клюйков / Л.П. Томкович Н.А. Кириллина [M.G. Pimenov, E.V. Klyuykov / L.P. Tomkovich, N.A. Cyrillin] / 5 июня 1977 [5 June 1977]" (https://plant.depo.msu.ru/open/public/item/MW0723305 - MW 0723305.jpg - https://www.gbif.org/occurrence/1697840793; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.); "Diphelypaea tournefortii, Berdatak, 2010.05.18 // Tsaghkadzor [Tsakhkadzor / Цахкадзор / Darachichag]" (photos 5788747528409755042 and 179759 [https://www.inaturalist.org/observations/80313] by Vahe Martirosyan [Ваэ Мартиросян] in Flora of Armenia's Check List and iNaturalist; accessed et rev. Ó. Sánchez Pedraja Feb 2013); "Цветок. Армения, область Котайк, окрестности Цахкадзора [neighborhood Tsakhkadzor] 31.05.2008" (photo 107214 by Ваэ Мартиросян [Vahe Martirosyan], 2011 in Plantarium [Diphelypaea tournefortii]; accessed Feb 2013); "Kotayk' [Yelgovan (Abovyan / Абовян, former Ellar district)], Armenia" (http://static.panoramio.com/photos/original/57543824.jpg by Vahe Martirosyan in www.panoramio.com, accessed Mar 2012); "Цветущее растение. Армения, обл. Котайк, окр. села Вохчаберд [env. Voghjaberd village], разнотравная степь [grassland steppe]. 26.05.2012" (http://www.plantarium.ru/page/image/id/166927.html et http://www.plantarium.ru/page/image/id/166926.html by Владислав Куропаткин [Vladislav Kuropatkin] in Plantarium, Diphelypaea tournefortii; accessed et rev. Ó. Sánchez Pedraja Feb 2013); "Diphelypaea tournefortii, Tsaghkadzor, 2014.06.04" (photo 6024440769631112354 by Vahe Martirosyan [Ваэ Мартиросян] in http://vahemart.livejournal.com; accessed et rev. Ó. Sánchez Pedraja Jan 2015); "Photo taken in Jrvezh Forest Park, H-3, Jrvezh [Джрвеж], Armenia 40° 10' 35.44" N  44° 37' 51.03" E" (http://static.panoramio.com/photos/large/3909676.jpg by Грант Веапетян in www.panoramio.com; accessed et rev. Ó. Sánchez Pedraja Jan 2015, sub P. tournefortii); "Armenia, near Tseghkadrov city, dry grassland 40,524303º N ; 44722640º E. [40°31'27.5"N 44°43'21.5"E 40.524303, 44.722640] 15.05.2014. M. Peregrym" (Леонідович [Leonidovich] 2016: 106, sub P. tournefortii); "Tsakhkadzor [40°32'33.3"N 44°45'04.7"E 40.542570, 44.751310, near Hrazdan / Раздан], 5 Jun 2022" (https://www.inaturalist.org/observations/120286216 by https://www.inaturalist.org/people/ioghony, sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, stem and calyx with typical hairs, corolla-lobes overlapping).

Vayots Dzor Province / Вайоцдзорская область (Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii): "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армянская ССР, Даралaгез [Armenian SSR, Daralagez]. Гора Теке-Долдуран, урочище Xазма [T’ek’sari Lerrnashght’a range?, natural boundary Hazma?], / горные луг [mountain meadow], 10 июля 1929 г. [10.07.1929] / Собрали Ант. и И.llyкины [Leg. Ant. and I. llykin]" (https://plant.depo.msu.ru/open/public/item/MW0723304 - MW 0723304.jpg - https://www.gbif.org/occurrence/1697840810; rev. Ó. Sánchez Pedraja Apr 2018 [!]); 1. "... / Instituti Botanici Academiae Scientiarum / Armeniae / Phelypaea bayernii Novop. / Микояновский район [Mikoyan Region / Mikoyanovsky District (administrative unit from the Soviet era that is known today as the Yeghegnadzor / Municipality, formerly Keshishkend, within the Vayots Dzor Province)], ... / Karakaya [Каракая / Garagaya, today Sevazhayr / Севажайр (Yeghegis Community), 4.5 km SW of Getikvank / Gədikvəng / Гетикванк], calcareous... / Leg. Ш. Асланян [Sh. Aslanyan] ... / 15.VI.1957 Det. Ja. Mulkidjanjan". – 2. "Phelураea tournеfortii Desf. / Determ. ..." (https://data.rbge.org.uk/herb/E01283803 - E 01283803.zip - https://www.gbif.org/occurrence/4420620351; rev. Ó. Sánchez Pedraja Jul 2016, sub P. tournefortii (Piwowarczyk & al. 2019: 21, sub D. tournefortii)); "... / Diphelypaea tournefortii ... / ... Armenia, Vayotsdzor province, Vayk distr., side of volcano Vayots Sar, loc. Topasi / Dalik, 4 km SWW of village Karmrashen, 2400 m.s.m.; 39º48' / 45º29'E / Leg. M. Barkworth, F. Smith, E. Gabrielian, A. Nersesyan, / 27.06.2003 / Dt. M. Oganesian 2003" (UTC 00239126 - https://s.idigbio.org/idigbio-images-prod/3a60a4764186884f15b7485bbeabe6d3 - https://www.gbif.org/es/occurrence/1986951694; rev. Ó. Sánchez Pedraja Feb 2020, sub P. tournefortii); Proshaberd [39°49'54.1"N 45°22'27.0"E, near Vernashen, Gladzor Municipality], 18 Jun 2012 (https://www.inaturalist.org/observations/23155801 - https://www.gbif.org/occurrence/2823126326 by Christoph Schmidtlein, sub Phelypaea coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jun 2021, the photos correspond to P. tournefortii on Tanacetum sp.); "Цветки. Армения, Селимский перевал [Selim Pass / Orbelian's Caravanserai]. 20.05.2014" (http://www.plantarium.ru/page/image/id/238873.html by Тамара Галстян [Tamara Galstyan] 2014 in Plantarium, sub Diphelypaea tournefortii; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [!]); "Цветок. Армения, Вайоц Дзор, окр. монастыря Спитакавор [Vayots Dzor, env. Spitakavor monastery]. 02.05.2013" (http://www.plantarium.ru/page/image/id/188448.html et http://www.plantarium.ru/page/image/id/188447.html by Марина Скотникова [Marina Skotnikova] 2013 in Plantarium, sub Diphelypaea tournefortii; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [!]); "Vayots Dzor Province [39°37'18.9"N 45°20'58.1"E 39.621920, 45.349480, near Boloraberd / Kurtkulag / Qurdqulaq], 11 Jun 2023" (https://www.inaturalist.org/observations/167549951 by https://www.inaturalist.org/people/karen3049 [Karen Olianti], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, stem and calyx with conspicuous hairs, apparently on Tanacetum); "Wajoz Dsor [39°47'35.8"N 45°29'32.9"E 39.793290, 45.492480, Herher, Jermuk Municipality], 29 Jun 2023" (https://www.inaturalist.org/observations/170225598 - https://www.gbif.org/occurrence/4145448144 by https://www.inaturalist.org/people/hyacinthus [Jacek Pietruszewski], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii); "Wajoz Dsor [39°47'41.2"N 45°29'35.2"E 39.794770, 45.493100, next to Mount Vayotssar / Dalik Tapa, Herher, Jermuk Municipality], 29 Jun 2023" (https://www.inaturalist.org/observations/170224213 - https://www.gbif.org/occurrence/4145400183 by https://www.inaturalist.org/people/hyacinthus [Jacek Pietruszewski], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, apparently on Tanacetum); "Vayots Dzor Province [39°49'23.1"N 45°08'53.9"E 39.823070, 45.148310, near Yelpin (Areni Municipality)], 3 May 2024" (https://www.inaturalist.org/observations/338226108 - https://www.gbif.org/es/occurrence/6159569715 by https://www.inaturalist.org/people/attila21 [Attila Steiner], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "Highway T-9-31 [39°37'31.0"N 45°34'54.4"E 39.625290, 45.581790, Gomk / Гомк (Vayk Municipality)], 13 May 2025" (https://www.inaturalist.org/observations/281271045 by https://www.inaturalist.org/people/goatnotes [Jonathan Tales], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, apparently on Tanacetum); "Vayots Dzor [39°48'51.2"N 45°38'44.9"E 39.814220, 45.645810, near Jermuk], 15 Jun 2025" (https://www.inaturalist.org/observations/362441884 - https://www.gbif.org/es/occurrence/6352631342 by https://www.inaturalist.org/people/vrufray [Vincent Rufray], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base, stem and calyx with conspicuous hairs).

Yerevan / Erevan / Erywan / Jerewan / Ереван (Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii): "Армения [40°10'30.8"N 44°37'14.8"E 40.175220, 44.620790, Jrvezh park, Yerevan], 16 May 2023" (https://www.inaturalist.org/observations/161975800 - https://www.gbif.org/occurrence/4120892826 by https://www.inaturalist.org/people/marinemeer, sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, plants with scales only at the base, stem and calyx with conspicuous hairs, apparently on Tanacetum); "Армения [40°10'35.1"N 44°37'15.8"E 40.176420, 44.621050, at Jrvezh park, Yerevan], 24 May 2025" (https://www.inaturalist.org/observations/283630682 by https://www.inaturalist.org/people/jorushka, sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, plants with scales only at the base, stem conspicuous hairs, apparently on Tanacetum).

Unknown location: "Velmi výraznými prvky jsou zástupci zárazovitých (Orobanchaceae), např. nepřehlédnutelná rostlina s jediným červeným květem Phelypaea coccinea (obr. 10), která parazituje na chrpách (Centaurea)." (Ekrtová & Ekrt 2012: 70, 72, photo 10; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [the photo seems to correspond to P. tournefortii]).

inicio / top of page

Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaïdjan (Fischer 1852: 107, [Armenié, Bushe], sub Anoplanthus Tournefortii; Boissier & Buhse 1860: 169, sub Anoplanthus Tournefortii; Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis; Stapf 1915: 288, sub Phelypaea orientalis, flore coccinea; Beck 1930: 44, sub P. tournefortii; Karjagin 1957)

Khizi District / Xızı rayonu / Хызинский район: 1. "Anoplon Bieber / steinii / ... 27 Juin 56 [1856?] // Armenia Rossica [Dübrar Dağı?, near Altıağac / Alty-Agatsch?] / Seidlitz [N.K. von Seidlitz] / Herb. Al. de Bunge". – 2. ibidem (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970939 - P 02970939.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/439109406; rev. Ó. Sánchez Pedraja Nov 2011, sub D. tournefortii [three (2+1) specimens on the lower part of the sheet, close to Bunge's labels]); "Russian Armenia" (Stapf 1915: 288, sub Phelypaea orientalis, flore coccinea); "Calculated" (Domina & Raab-Straube 2010, sub P. tournefortii).

Khojavend District / Xocavənd rayonu / Ходжавендский район [unconfirmed]: "Tale Dschagritschai, bei Takjalta Kulu [Dağdöşü / Tak] (Bayern!)" (Beck 1930: 44, sub P. tournefortii).

Nakhchivan Autonomous Republic / Naxçıvan Muxtar Respublikası / Нахичеванская Автономная Республика (Boissier & Buhse 1860: 169, sub Anoplanthus Tournefortii)

Babek District / Бабекский район / Babək rayonu: 1. "444. Anoplanthus Tournefortii / Walp. ex parte / Puskoh bei Nachitschewan [Nakhchivan / Naxçıvan / Нахичевань] / 31.V, 47 [31 May 1847] / Buhse [leg.] // Iter Persicum. / Buhse, 1847-1849 / Herb. Al. de Bunge". – 2. "Phelipaea coccinea in MNHN-P-P02970936" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970936 - P 02970936.jpg - E. Cosson - https://www.gbif.org/occurrence/438443080; rev. Ó. Sánchez Pedraja Nov 2011, sub P. tournefortii [the two specimens on the upper left corner close to Buhse's label]); "Phelypaea coccinea (M. Bieb.) Poir. / Nakhichivan [Nachitschewan / Naxçıvan / Нахичевань] / 1847 / F. Buhse" (LD:General:2073901 - https://www.gbif.org/es/occurrence/2012691834 [n.v.]); "Bei Nachitschewan im Dschagrithal [Dzhagry / Cəhriçay / Джагричай, Babek District / Бабек] beim Dorfe Pusko, auf Pyrethrum achillaefolium (nicht auf P. myriophillum, wie Fischer Bull de la Soc. Imp. de Moscou 1852 nº I 107, angiebt), 31 Mai 1847 (florens). Nº 444" (Boissier & Buhse 1860: 169, sub Anoplanthus Tournefortii); "Armeniâ Rossicá (Buhse!)" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Transcaucasia: Near Pusko in the Dshagri valley [Dzhagry / Cəhriçay], north of Nahitshewan [Nachitschewan / Nakhchivan / Нахичевань], on Pyrethrum myriophyllum, C.A. Mey., Buhse" (Stapf 1915: 293, sub P. tournefortii); "in der Provinz Nachitschevan [Nakhchivan Autonomous Republic] um Nachitschevan [Nakhchivan / Naxçıvan] bei Puskoh (Buhse!)" (Beck 1930: 44, sub P. tournefortii); "Herbarium Universitatis Mosquеnsis (MW) / ... / Filiatonis Azerbajdzhanicae Academiae Scientiarum URSS / Herbarium Azerbajdzhanicum / Phelураea coccinea Poir. / Transcaucasia, Azerbajdzhan, respublica Nachitshevan, distr. / Nachitshevan [Nachitschewan / Naxçıvan / Нахичевань], prope pag. Buzgov [Aşağı Buzqov]. / in calcareis 29.V.1939 Leg. E. Chalivov / Det. ..." (https://plant.depo.msu.ru/open/public/item/MW0723302 - MW 0723302.jpg - https://www.gbif.org/occurrence/1697840776; rev. Ó. Sánchez Pedraja Apr 2018, sub P. tournefortii).

Ordubad District / Ордубадский район / Ordubad rayonu: 1. "Herbarium Instituti botanici nom. V.L. Komarovii Ac. Sc. URSS / Азербайджан Нахичеванская ССР [Nakhichevan Autonomous Soviet Socialist Republic / Nakhichevan District [an old name of Babek District], Anabad [Анабад] locality on the [Карагут (Каракий)] Karagut / Karakut / Karakush (Karaksiy Mount Kaputjugh / Qapıcıq / Капутджухh (Zangezur range / Зангезурский хребет)]) massif, approximately 1600 m on the northern grassy slopes // leg. A.A. Grossheim, I.A. Ilyinskaya and M.I. Kirpichnikov / 9.VI.1947] ...". – 2. "Phelypaea Tournefortii / Desf. / 6.XI.1947 / Determ. I. Novopokrovsky". – 3. "Anoplon Tournefortii (Desf.) G. Don / 1966 rev. A. Gilli" (https://data.rbge.org.uk/herb/E01283802 - E 01283802.zip - https://www.gbif.org/occurrence/4420521875; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii).

Sadarak District / Садаракский район / Sədərək rayonu: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / В 2 км к югу от с. Кярки [2 km to the south of the village Karki / Kərki / Kyarki / Tigranashen / Тигранашен] можжевеловое редколесье, 1400 м. / 29 мая 1973 г. [29.05.1973] Н. Шведчикова [N. Shvedchikova]" (https://plant.depo.msu.ru/open/public/item/MW0723306 - MW 0723306.jpg - https://www.gbif.org/es/occurrence/1697840797; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.).

Sharur District / Шарурский район / Şərur rayonu: 1. "Herbarium Universitatis Mosquеnsis (MW) / ... / ... Зривано-Нахичеванская ... / Phelураea coccinea Poir. / Нахреспублика [Nakhrespublika / Nakhchivan Autonomous Republic], хребет Сарайбулаг [Saraybulag ridge / Eranos Range (Armenian Highlands [Sadarak District-Sharur District])] / 10.VI.1931 / Leg. И.И. Карягин [I.I. Karyagin] Det. A.A. Grossheim". – 2. "Diphelypaea coccinea in MW" (https://plant.depo.msu.ru/open/public/item/MW0723298 - MW 0723298.jpg - https://www.gbif.org/occurrence/1697840765; rev. Ó. Sánchez Pedraja Apr 2018, sub D. tournеfortii [P. tournefortii]); "Herbarium Universitatis Mosquеnsis / Azərbaycan Mərkəzi Herвarium /... / Phelураea Tournеfortii Desf. / Transcaucasia, respublica Naсhitschevan, prope p. / Karakush, in herbosis. / Leg L. Prilipkо / 31-V-1934 Det. L. Prilipkо" (https://plant.depo.msu.ru/open/public/item/MW0723308 - MW 0723308.jpg - https://www.gbif.org/occurrence/1697840815; rev. Ó. Sánchez Pedraja Apr 2018); 1. "Herbarium Instituti botanici nom. V.L. Komarovii Ac. Sc. URSS / Азербайджан Нахичеванская ССР [Nakhichevan Autonomous Soviet Socialist Republic / Nakhichevan District [an old name of Babek District], near the Alma-bulag (Ağbulaq / Ağbulaq Yolu, near Yuxarı Yayci) spring ... 1900 m ... subalpine ... / leg. A.A. Grossheim, I.A. Ilyinskaya and M.I. Kirpichnikov / 10.VI.1947]". – 2. "Phelypaea Tournefortii / Desf. / 6.XI.1947 / Determ. I. Novopokrovsky". – 3. "Anoplon Tournefortii (Desf.) G. Don / 1966 rev. A. Gilli" (https://data.rbge.org.uk/herb/E01283808 - E 01283808.zip - https://www.gbif.org/occurrence/4420516370; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii).

inicio / top of page

Georgia / Sakartvelo / Georgien / Géorgie / Грузия [needs be confirmed] (Piwowarczyk & al. 2023: 14, sub P. coccinea)

"Anoplon Biebersteinii C.A.M. / Caucas / 1850. / leg. Kovalevsky" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970929 - P 02970929.jpg - hb. Drake - https://www.gbif.org/occurrence/439442856; rev. Ó. Sánchez Pedraja Nov 2011, sub P. tournеfortii, only one specimen well preserved, "calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes; scale-leaves concentrated towards the bases of the stems" (Cullen 2010: 384)); "Georgia" (Tarkhnishvili & Chaladze 2013 in Georgian Biodiversity Database, accessed Dec 2010 and Oct 2017); "Fl. rossica / Anoplanthus Biebersteinii Reut. / Tiflis / leg Bienert [Theophil Joachim Heinrich Bienert, c. 1859]" (https://www.botanicalcollections.be/#/en/specimen/BR0000031258340 - https://oxalis.br.fgov.be/images/BR0/000/031/258/340/BR0000031258340.jpg - BR 0000031258340.jpg - https://www.gbif.org/occurrence/4072994557; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii [the three shorter plants on the left side of the sheet with hairy calyx and scales at the base, which apparently parasitizing on Tanacetum] and P. coccinea [only the longest plant on the right side of the sheet with the scales along the stem]. P. tournefortii is not yet known from Georgia; perhaps the sheet contains two collections from different places).

Note: The gathering of Hohenaker in May 1831 and 1834 seem to correspond to D. coccinea: "pratis montosis umbrosis Georgiae caucasicae (Hohenacker !)" (Reuter 1847: 42, sub Anoplanthus biebersteinii Reut.); "in pratis montosis Georgiae Caucasicae (Hoh!)" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Georgien (Hohenacker nach Boissier)" (Beck 1930: 44, sub P. tournefortii).

inicio / top of page

Iran / ایران‎ / Irán [Persia] [see also P. boissieri] (Gardner & Gardner 2014: 119, sub D. tournefortii)

Kurdistan Province [unconfirmed/n.v.]: "Planta of the Orient, 1963; May 14. M. Jacobs 6573 / ... / Phelypaea coccinea / Northern and western Iran. Place 35º30'N 46º38'E [near Kaku Zakaria, Marivan County] / Region: West central Kordestan. Alt. 1600-1900 m. / ..." (https://data.biodiversitydata.nl/naturalis/specimen/L.2821535 - L 2821535.jpg - https://www.gbif.org/occurrence/2514613659 - https://www.europeana.eu/es/item/11651/_Botany_L_2821535; rev. Ó. Sánchez Pedraja Oct 2017, sub cf. P. boissieri); idem (https://data.rbge.org.uk/herb/E01283826 - E 01283826.zip - https://www.gbif.org/occurrence/4420619361; rev. Ó. Sánchez Pedraja Jul 2026, sub cf. P. boissieri); 1, idem – 2. "Phelypaea tournefortii in Kew" (http://specimens.kew.org/herbarium/K004381258 - https://records.data.kew.org/occurrences/b297cd9d-b89e-4713-82e6-bb4d8ae3738a - K 004381258.jpg - https://www.gbif.org/occurrence/5060753009; rev. Ó. Sánchez Pedraja Jul 2026, sub cf. P. boissieri).

West Azerbaijan Province: "Flora of Iran / Phelipaea toumefortii Desf. / Prov. Azerbaijan; E. of Bezorgan [Bazargan, Maku County]; 6000 ft. / alt.; on dry rocks at foot of cliff. / steem deep crimsom; bracts bright crimson; anthers projecting; pale yellow; / stem hairy; no leaves. / Furse & Synge 74. 22.5.1960" (http://specimens.kew.org/herbarium/K004381261 - https://www.gbif.org/occurrence/5060755990 - K 004381261,jpg - https://records.data.kew.org/occurrences/2adaf8b4-d780-4352-a22d-6222a942541a; rev. Ó. Sánchez Pedraja Jun 2021, sub P. tournefortii, plants with scales only at the base of the hairy stem and calyx with conspicuous hairs); "Rezaiyeh College of Agriculture / and Animal Husbandry [Urmia University] / No. 5207 / Phelipaea toumefortii Desf. / (larger specimen ...) / W. Azerbaijan: Maku / 1200 m / 11.5.73 / Coll. Siami" (http://specimens.kew.org/herbarium/K004381265 - K 004381265.jpg - https://records.data.kew.org/occurrences/2bc99d95-2032-4ccf-a4fc-a94669a44a8a - https://www.gbif.org/occurrence/5060747982; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); 1. "K.H. Rechinger, Iter Orientale 1971 / Flora of Persia / Phelipaea coccinea / Prov. Azerbaijan [West Azerbaijan]: Ghogeh Dagh [Gecheh Dāgh (Marakan Protected Area, Armenian highlands)], near Bazargan [Maku County (Turkish frontier), Āzārbāyjān-e Gharbī]. / limestone slopes and rocks. / Alt. c. 2200-2250 m  1 august 1971 / Coll. Jennifer Lamond No. 5000". – 2. "Herb. Musei Hist. Natur. Vindob. [Naturhistorisches Museum Wien] / Anoplon coccineum (M. B.) H. Riedl et / Schiman-Czeika / 1973 rev. A. Gilli"  (https://data.rbge.org.uk/herb/E01283825 - E 01283825.zip - https://www.gbif.org/occurrence/4420620353; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base, stem and calyx with conspicuous hairs and apparently on Tanacetum sp.); "W: Azerb. [West Azerbaijan]: Ghogeh Dagh [Gecheh Dāgh (Marakan Protected Area, Armenian highlands)] prope Bazorgan [Bazargan, Maku County (Turkish frontier), Āzārbāyjān-e Gharbī], 2200.2250 m, Lamond 5000." (Gilli 1979: 290, sub Anoplon coccineum [rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii]); "Urmia region?" (https://gramhir.com/media/2587342286867981834 by @hadi.urmulu [https://gramhir.com/profile/hadi.urmulu/2303903103], sub P. coccinea; accessed et rev. Ó. Sánchez Pedraja Sep 2021 [sub P. tournefortii, apparently on Tanacetum sp.]).

inicio / top of page

Turkey / Türkiye / Tuquía / Turkije (Reuter 1847: 43, sub Anoplanthus tournefortii; Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis; Stapf 1915: 293, sub P. tournefortii; Beck 1930: 44, sub P. tournefortii; Gilli 1982: 2, sub P. tournefortii; Bakis & al. Distribution of The Taxon Over Turkey in Turkish Plants Data Service (TÜBİVES) (accessed Sep 2017); Bizim Bitkiler 2021 (http://www.bizimbitkiler.org.tr...id=9705, sub P. tournefortii); Piwowarczyk & al. 2019: 18, sub D. tournefortii, epitype of P. tournefortii)

Central East Anatolia Region / Ortadoğu Anadolu Bölgesi

Malatya Province / Malatya ili: "Malatya dolaylarında çekilmiştir [it was taken near Malatya]" (photos http://yabanicicekler.com/flower/phelypaea-coccinea-53 [1/3-2/3-3/3 (53)] sub Phelypaea coccinea in Türkiye'nin Çiçekleri - Yabani Çiçekler; accessed et rev. Ó. Sánchez Pedraja Oct 2017 [the three photos seem to correspond to P. tournefortii ("Stems and leaves covered with dense to sparse, long, multicellular white hairs; calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes" (Cullen 2010: 384)), moreover we can observe the host-plant [Tanacetum sp.] in the photo 2/3]).

Tunceli Province / Tunceli ili [formerly Dersim Province]: "Flora Turcomanniae Asiaticae / Iter Leydense 1959 / Collectores: E. Hennipman, P. Nijhoff, C. Swennen, A.S.Tulp, / W.J.M. Vader, V.J.J.O. de Wildee / No. 1635 Date: 29-5 Altitude ± : 2000 m. / Prov.: Tunçeli. Locality: ± 5 km N. of Pülümür (along the road Elâzig-Erzincan) / ... / Flowers humid neglected field. Clayish soil. / Flowers bright red. / (Host Centaurea ... / Anoplanthus coccinus Walp." (https://data.biodiversitydata.nl/naturalis/specimen/L.2825068 - L 2825068.jpg - https://www.gbif.org/occurrence/2514627751; rev. Ó. Sánchez Pedraja Jun 2021, sub P. tournefortii, plants with scales only at the base of the hairy stem and calyx with conspicuous hairs; the indicated host-plant (Centaurea) is apparently erroneous but see P. boissieri); idem (https://data.biodiversitydata.nl/naturalis/specimen/WAG.1141086 - WAG 1141086.jpg - https://www.gbif.org/occurrence/2515854856; rev. Ó. Sánchez Pedraja Jun 2021, sub P. tournefortii but see P. boissieri); idem (http://specimens.kew.org/herbarium/K004381234 - K 004381234.jpg - https://records.data.kew.org/occurrences/291c8caa-8145-4248-a3b5-89079b101b65 - https://www.gbif.org/occurrence/5060755987; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base and calyx with conspicuous hairs); "Tunceli, S¸Y 3245" (Zare & al. 2014, sub D. tournefortii); "Karagöz, Pülümür [39°26'49.8"N 39°40'05.0"E], 2 jul 2015" (https://www.inaturalist.org/observations/38100719 - https://www.gbif.org/es/occurrence/2563645191 by Mustafa Gökmen, sub Diphelypaea coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Feb 2020, sub P. tournefortii, apparently on Tanacetum sp.).

Van Province / Van ili: "Coteaux secs, entre Van et Hokotz-vank ['Hokvotz-vank, monastére isolé, á environ 45 kilom. au sud-sudest de Van, tout près et au sud-ouest de Soussouz-dagh'] (exs. 110)" (Aznavour 1917: 28, sub Anoplanthus coccineus var. peduncularis Boiss.); "zwischen Van und Hokotz-vank (D. Tchitouny nach Aznavour 1917)" (Beck 1930: 44, sub P. tournefortii); "Van: between Van and Hokotz church, Tchitouny" (Gilli 1982: 2, sub P. tournefortii); "Sommerwurz in der Osttürkei nahe der iranischen Grenze, June 1990" (https://commons.wikimedia.org/wiki/File:Phelypaea_tournefortii_1.jpg by Ziegler175; accessed Feb 2020); "Bild 117 - Phelypaea toumefortii (Orobanchaceae) auf Tanacetum schmarotzend, am Fuß eines felsigen Hanges nördlich von Muradiye; etwa natürliche Größe." (Sorger 1994: 174, sub P. tournefortii); "June 13, 2002. Near Caldiran [Çaldıran, Van], eastern Turkey" (sub P. coccinea, 2006 Old Dominion University, accessed Dec 2010); "Van: steppes le long du chemin d’accès aux chutes de Muradiye, 1826 m, UTM 38S 0392562 / 4324169, leg. P. Coulot & al., 24.05.2009" [habit, flower, calyx, scale-leaves]; Lake Van / Phelypaea tournefortii - parasite on Tanacetum species (http://www.fands.org.uk/FANDS_images/VAN/_0012090.jpg in http://www.fands.org.uk; accessed Feb 2013); "Van" (https://turkiyebitkileri.com/foto.../phelypaea-tournefortii.html, sub P. tournefortii, in https://turkiyebitkileri.com/ [Plants of Turkey]; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "Tuşba [Tushpa / Tushpa-Van, 38°46'24.0"N 43°29'21.8"E 38.773320, 43.489400, near Meydancık] / between Van and Muradiye / 24 May 2006" (https://www.inaturalist.org/observations/339789798 - https://www.gbif.org/occurrence/6163188002 by https://www.inaturalist.org/people/roger_e_n_smith, sub P. tournefortii, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, apparently on Tanacetum sp.); "Muradiye [39°04'24.7"N 43°44'46.7"E 39.073528, 43.746306], 30 May 2015" (https://www.inaturalist.org/observations/338904479 - https://www.gbif.org/occurrence/6159135355 by https://www.inaturalist.org/people/langli [Jens Thalund], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, apparently on Tanacetum sp.).

East Black Sea Region / Doğu Karadeniz Bölgesi

Gümüşhane Province / Gümüşhane ili: "Tobey 2047 / Phelipaea coccinea / ? (Parasite) / (on Tanacetum) / Turkey. A7 Gümüşhane: Kep Köyu [Kov Kalesi?], 1300 m. / open / rocky slope ... 19 June 1967". – 2. "Phelipaea tournefortii Desf. / Det. J.R. Edmondson 10.4.77" (https://data.rbge.org.uk/herb/E01283812 - E 01283812.zip - https://www.gbif.org/occurrence/4420518388; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii).

Eastern Anatolia Region / Doğu Anadolu Bölgesi "Phelypaea tournefortii: IMG_3551, IMG_3555, IMG_3579 [on Tanacetum sp.] (photos by Dana Cromie in Eastern Turkey, May 2011 [https://www.flickr.com]; accessed et rev. Ó. Sánchez Pedraja Feb 2013)

Erzincan Province  / Erzincan ili: "Erzincan Merkez/Erzincan [formerly Yerznka, 39°33'51.1"N 39°45'20.2"E 39.564190, 39.755620, near Kalecik], Turecko, 16 Jun 1993" (https://www.inaturalist.org/observations/36848776 - https://www.gbif.org/occurrence/3986004340 by https://www.inaturalist.org/people/mnauky, sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base).

Hakkâri Province / Hakkâri ili: "Keçili Köyü, Yüksekova [37°38'51.9"N 44°17'32.7"E 37.647750, 44.292420], 21 May 2021" (https://www.inaturalist.org/observations/130141003 - https://www.gbif.org/occurrence/3892594576 by https://www.inaturalist.org/people/mervetugce, sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base of the hairy stem).

Iğdır Province / Iğdır ili: 1. "Plantae prov. Erivan (Armenia rossica) / Anoplanthus coccineus Wal. / Эриванская губ. [Erivan Governorate / Эриванская губерния] Сурмалинский у. [Surmalinskiy uyezd / Surmalu uezd (the district made up most of the Iğdır Province of present-day Turkey)], близ с. Сичанлу [near the village of Sürmeli (Tuzluca District)]. 14.VII". – 2. "Ex Herb. Inst. Bot. Ac. Sc. URSS.". – 3. "Diphelipaea coccinea in MW". – 4. "Diphelypaea coccinea (M.Bieb.) Nicolson in Catalogue of Life" (https://plant.depo.msu.ru/open/public/item/MW1020073 - MW 1020073.jpg - https://plant.depo.msu.ru/open/public/item/MW1020073...e3189153-acc2-4481-9960-a1b335e54eaf - https://www.gbif.org/occurrence/5888738139; rev. Ó. Sánchez Pedraja Aug 2026, sub  P. tournefortii, plants with scales only at the base, stem and calyx with conspicuous hairs).

Northeast Anatolia Region / Kuzeydoğu Bölgesi

Ağrı Province / Ağrı ili: "Davis 44038 / Phelipaea tournefortii Desf. / cited Fl. Turk. / Turkey. B9; 3-5 km from / Tutak to Hamur (Murat valley). / 1600 m / Sloping meadows and / rocky slopes. / Parasite on / Tanacetum species [= Tanacetum aureum (Lam.) Greuter, M. V. Agab. & Wagenitz (?)]. flowers scarlet with 2 black bosses / at throat! / 2 june 1966.". – 2. "Herb. Musei Hist. Natur. Vindob. / Anoplon tournefortii (Desf.) G. Don / 1966 rev. A. Gilli" (https://data.rbge.org.uk/herb/E01283809 - E 01283809.zip - https://www.gbif.org/occurrence/4420518387; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii); "Davis 44038 / Phelipaea tournefortii Desf. / cited Fl. Turk. / Turkey. B9; 3-5 km from / Tutak to Hamur (Murat valley). / 1600 m / Sloping meadows and / rocky slopes. / Parasite on / Tanacetum species [= Tanacetum aureum (Lam.) Greuter, M. V. Agab. & Wagenitz (?)]. flowers scarlet with 2 black bosses / at throat! / 2 june 1966." (http://specimens.kew.org/herbarium/K004381263 - K 004381263.jpg - https://records.data.kew.org/occurrences/6d9886bc-9984-4b12-aa8c-3e5fd2ed1a3a - https://www.gbif.org/occurrence/50607569801; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii); 1. "Herb. Hort. Bot. Reg. Edin. / Cambridge Botanical Expedition / Flora of Turkey / Phelypaea boissiera / Parasitic on Achillea sp. [= Tanacetum aureum ?] vilayet / Agri [Ağrı Province] Tahirdag [Tahir Dağı (Tahir Dag pass / Mount Ararat / Ağrı Dağı region)]. Grassy slopes. / Alt. c. 2600 m 13 July 1968 / Coll. E.M. Rix et al. No. 840". – 2. "Phelypaea boissiera Stapf. in E". – 3. "Diphelypaea coccinea var. boissieri (Reut.) Uhlich in Catalogue of Life"  (https://data.rbge.org.uk/herb/E01283801 - E 01283801.zip - https://www.gbif.org/occurrence/4420518386; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "B9 Ağri: Murat valley, 1600 m, D. 44038!" (Gilli 1982: 2, sub P. tournefortii); "Doğubeyazıt" (https://turkiyebitkileri.com...phelypaea-tournefortii/do%C4%9Fubeyaz%C4%B1t.html, sub P. tournefortii, in https://turkiyebitkileri.com/ [Plants of Turkey]; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "Doğubayazıt [39°29'52.6"N 44°00'04.5"E 39.497930, 44.001250, near Ortadirek], 13 Jun 2002" (https://www.inaturalist.org/observations/106223743 - https://www.gbif.org/occurrence/4507795570 by https://www.inaturalist.org/people/barbaraparris [Barbara Parris], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, apparently on Tanacetum sp.).

Bayburt Province / Bayburt ili: 1. "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia". – 2. "Phelipaea coccinea in MNHN-P-P02970936" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970936 - P 02970936.jpg - E. Cosson - https://www.gbif.org/occurrence/438443080; rev. Ó. Sánchez Pedraja Nov 2011, sub P. tournefortii [only one specimen on the lover left corner close to Huet de Pavillon's label]); 1. "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia". – 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855". – 3. "Phelipaea coccinea in MNHN-P-P02970941" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970941 - P 02970941.jpg - https://www.gbif.org/occurrence/438443082; rev. Ó. Sánchez Pedraja Nov 2011, epitype of P. tournefortii (Piwowarczyk & al. 2019: 18, sub D. tournefortii)); idem (http://specimens.kew.org/herbarium/K004381269 - K 004381269.jpg - https://records.data.kew.org/occurrences/24603291-387d-45ce-a545-46fdb686452a - https://www.gbif.org/occurrence/5060755064; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii [two small specimens close to their label]); "Bayburt" (Distribution of The Taxon Over Turkey in Turkish Plants Data Service (TÜBİVES) Version 2.0 BETA; accessed Sep 2017).

Erzurum Province [Western Armenia]: "Armenia [Turkish Armenia / Armeniâ Turcicâ / Western Armenia]" (Desfontaines 1807: t. 21; 1808: 16, t. 10); 1. "Anoplanthus / Phelipaea Tournefortii Desf. / Ex ... Tournefortii / Choix de Plantes du Corollaire". – 2. "Phelipaea coccinea in MNHN-P-P02970942" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02970942 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970942 - P 02970942.jpg - https://www.gbif.org/occurrence/437111828; rev. Ó. Sánchez Pedraja Jun 2021, sub P. tournefortii); "H. Native of Armenia [Turkish Armenia / Western Armenia]" (Don 1838: 633, sub Anoplon tournefortii); 1. "Kurdistan [Turkish Kurdistan] / J. Brant, W.H.F. Strangways, 1840 [Herbarium Benthamianum 1854]". – 2. "Host: Pyrethrum / myriophyllum / C.A.M.". – 3. "Anoplanthus Tournefortii / Walp . ex / Reut.". – 4. "Phelipaea Tournefortii Desf. / Determinavit O.S. [O. Stapf]" (http://specimens.kew.org/herbarium/K004381267 - K 004381267.jpg - https://records.data.kew.org/occurrences/74032894-2aab-43a1-9ca5-5e7dabbcb4bf - https://www.gbif.org/occurrence/5060753010; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "In Armeniâ [Turkish Armenia / Western Armenia] (Tournef.)" (Reuter 1847: 43, sub Anoplanthus tournefortii); 1. "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia". – 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie [Turkish Armenia / Western Armenia]. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855". – 3. "Phelipaea coccinea in MNHN-P-P02970941" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02970941 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970941 - https://mediaphoto.mnhn.fr/media/1441352983821Tbv218qAE1INSMiF - P 02970941.jpg - https://www.gbif.org/occurrence/438443082; rev. Ó. Sánchez Pedraja Nov 2011, sub P. tournefortii); 1. "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia". – 2. "Herbarium Kewense. / Ex legato alexander Prior. / Mense Aprili, 1903". – 3. "Phelipaea / coccinea / Marsch [handwritten on the sheet]". – 3. "Orobanche coccinea subsp. coccinea in Kew". – 4. "Diphelypaea coccinea (M.Bieb.) Nicolson in Catalogue of Life" (http://specimens.kew.org/Herbarium/K004381252 - K004381252.jpg - https://records.data.kew.org/occurrences/3ed7d085-1dd2-4a25-8e5e-a836b71a6727 - https://www.gbif.org/occurrence/5060755989; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii, plants with scales only at the base); 1. "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia". – 2. "Phelipaea coccinea in MNHN-P-P02970936" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970936 - P 02970936.jpg - E. Cosson - https://www.gbif.org/occurrence/438443080, rev. Ó. Sánchez Pedraja Nov 2011, sub P. tournefortii [only one specimen on the lover left corner close to Huet de Pavillon's label]); 1. "Anoplon Tournefortii / ou / Phelipaea coccinea". – 2. "55. Top Dagh [Top Dağı (hill located 10 km northeast of Erzurum city, see Aziziye Tabyası)] près / Erzeroum [Erzurum], Arménie. / Simon, 1854" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970940 - P 02970940.jpg - E. Cosson - https://www.gbif.org/occurrence/437444715, rev. Ó. Sánchez Pedraja Nov 2011 [seven specimes well preserved]); 1. "Orobanchea / Anoplon Tournefortii / Syn.:? / Phelipaea coccinea?". – 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie [Erzurum?]. / Collection de Mr. Simon. [1854] / donnée par M^r. le C^te. de Challaye 1855". – 3. "Phelipaea coccinea in MNHN-P-P02970943" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02970943 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970943 - P 02970943.jpg - https://www.gbif.org/occurrence/438443081; rev. Ó. Sánchez Pedraja Nov 2011, sub P. tournefortii); 1. "Erzeroum [Erzurum] J. Zohrab". – 2. "Phelipaea Tournefortii Desf. / Determinavit O.S. [O. Stapf]" (http://specimens.kew.org/herbarium/K004381271 - K 004381271.jpg - https://records.data.kew.org/occurrences/a1d69d31-29e4-41fa-bd74-43f9fa305207 - https://www.gbif.org/occurrence/5060753999; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii [only one specimen]); "Anoplanthus Tournefortii / Armenia [Turkish Armenia / Western Armenia] Received May 1867 / 158 Coll. Messrs. H.H. Calver and J.Zohrab" (http://specimens.kew.org/herbarium/K004381270 - K 004381270.jpg - https://records.data.kew.org/occurrences/98a518ee-4e82-4583-b669-e8036ec5fae8 - https://www.gbif.org/occurrence/5060755991; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii [three specimens]); 1. "Anoplanthus Tournefortii / Armenia [Turkish Armenia / Western Armenia] Received May 1867 / 158 Coll. Messrs. H.H. Calver and J.Zohrab". – 2. "... Anoplanthus tournefortii (Desf.) G. Don / 1966 rev. A. Gilli" (https://data.rbge.org.uk/herb/E01283807 - E 01283807.zip - https://www.gbif.org/occurrence/4420644358; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "In Kurdistaniâ [Turkish Kurdistan] (J. Brant! [http://specimens.kew.org/herbarium/K004381267] Strangways in h. Benth.!)" (Reuter 1847: 43, sub Anoplanthus tournefortii); "Kurdistaniâ [Turkish Kurdistan] (Barré de Lancy [vice-consul de France à Mersine et Adana])!" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, terra class. of P. tournefortii Desf.] (Tourn!, Huet!, Calv!)" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Herbier de Ch. d'Alleizette / Plantes ... levant d'échanges [?] / Phelypaea coccinea Pers. / (Anoplanthus Tournefortii) / Asie M^re [Mineure] env. Erzurum / Juillet 1904" (https://data.biodiversitydata.nl/naturalis/specimen/L.2821536 - L 2821536.jpg - https://www.gbif.org/es/occurrence/2516614582; rev. Ó. Sánchez Pedraja Oct 2017); "discovered in Armenia, probably in the neighbourhood of Erzurum by Tournefort in 1701 [June 15 to July 6]" (Stapf 1915: 285, 289, 293, sub Phelypaea orientalis, flore coccinea and sub P. tournefortii); "native of Armenia [Turkish Armenia / Western Armenia]" (Stapf 1915: 285, 288, sub Phelypaea orientalis, flore coccinea); "Turkish Kurdistan" (Stapf 1915: 288, sub Phelypaea orientalis, flore coccinea); "Turkish Armenia: Upper Chorok valley, near Masut [Çakılı], E. of Baibut [Baiburt / Bayburt], 1200-1800 m, Huet de Pavillon! Near Erzurum, Zohrab, 826! [http://specimens.kew.org/herbarium/K004381271] Calver and Zohrab, 158! [http://specimens.kew.org/herbarium/K004381270] Kurdistan; without precise locality, Brant! [http://specimens.kew.org/herbarium/K004381267] Barre de Lancy (Hb. Gaillardot, 2555)" (Stapf 1915: 293, sub P. tournefortii); "ad radices Pyrethri myriophylli C.A. Meyer [= Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech (K 004381267)] et Achilleae spec. [= Tanacetum?] ... Armenien: Provinz Erzerum [Erzurum] (Tournefort, Calvert nach Boissier); im Tale Kussuklu zwischen Baiburt und Erzerum bei 4000-5000' (Huet de Pavillon!) ... Kurdistan (Barre de Lancy nach Boissier)" (Beck 1930: 44, sub P. tournefortii); "Flora of Turkey / Phelipaea tournefortii Desf. cited Fl. Turk. Sari Koms Pass / 8000 ft.  [2600 m] alt.; Alpine meadows / probably parasitic on Achillea [= Tanacetum?]. / Notes: 4-8'' tall. Stems maroon. No leaves. / single flowers, vivid scarlet with black / blotch in throat, shaped like an / .... / Anthers yellow / [Collector:] Furse & Synge 850 / 2.7.1960" (http://specimens.kew.org/herbarium/K004381262 - K 004381262.jpg - https://records.data.kew.org/occurrences/ca4725e1-11bd-4216-971b-40cf2028e19e - https://www.gbif.org/occurrence/5060753998; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii); 1. "Flora of Turkey / Phelipaea tournefortii Desf. / Prov. Armenia, Distr. Erzerum [Erzurum] / Sari Koms Pass, Alpine meadows / probably parasitic on Achillea / 2.7.1960  8000 ft. [2600 m] / Notes: 4-8'' tall ... / Collector: Furse & Synge 850". – 2. "... Anoplanthus tournefortii (Desf.) G. Don / 1966 rev. A. Gilli" (https://data.rbge.org.uk/herb/E01283810 - E 01283810.zip - https://www.gbif.org/occurrence/4420644359; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "Flora of Turkey / B8 Erzurum [Erzurum Province]; Hamurpet Daglari [Akdoğan Mountains]. From Karaağil, / to E[rzurum]. Mt. steppe: open, rocky / ... / 20 June 1967 / A.R. Mitchell  C.M. & W 2778". – 2. "Phelypaea coccinea (M. Bieb.) Poir. / Determinavit J.R. Edmondson 6.7.77" (https://data.rbge.org.uk/herb/E01283817 - E 01283817.zip; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base); 1. idem. – 2. "Phelipaea tournefortii Desf. / Turkey / A.R. Mitchell 2778" (http://specimens.kew.org/herbarium/K001513716 - https://records.data.kew.org/occurrences/aff634c1-226f-4480-b46d-edf93079de99 - K 001513716.jpg - https://www.gbif.org/occurrence/4519430108; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); 1. "Tobey 2047A / Phelipaea coccinea / Parasite / Turkey. A8 Erzerum [Erzurum]; Aras river Gorge [Mescitli Boğazı (Köprüköy district)]. 1650 m. / Bright scarlet when fresh. host compositae. / 20 June 1967". – 2. "Phelypaea tournefortii Desf. / Determinavit J.R. Edmondson 10.4.77" (https://data.rbge.org.uk/herb/E01283813 - E 01283813.zip - https://www.gbif.org/occurrence/4420521876; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); "B8 Erzurum [Erzurum Province]: Karaağil to Erzurum, Hamur-pet Da. [Akdoğan Mountains], Mitchell 2778(!)" (Gilli 1982: 2, sub Phelypaea coccinea)."E. Anatolia. A8 Erzurum: Sari Koma pass, 2600 m, Furse & Synge 850 [https://data.rbge.org.uk/herb/E01283810]! Gümüşane: Kussuklu valley, Bayburt to Erzurum, 1220-1525 m, Huet." (Gilli 1982: 2, sub P. tournefortii); "https://www.ncbi.nlm.nih.gov/nuccore/AY209285: Turkey, NE Anatolia, between Bayburt and Erzurum, between the village Maden and Kop Geçidi, 1840 m, N 40° 04' 49'' E 40° 25' 34'', on Tanacetum sp. (Asteraceae), G. Schneeweiss & C.-G. Jang (WU: S&T 7764 [http://www.parasiticplants.siu.edu/Orobanchaceae/images/Diphelypaea_tourn_habit.jpg]) (Carlón & al. 2005: 63 sub P. tournefortii Desf.)"; "Gümüşane, Mescit Daglari: an der Straße zwischen Bayburt und Erzurum, zwischen dem Ort Maden und dem (Pass) Kop Geçidi [Bayburt Province]; 40°04'49'' N, 40°25'34'' E. 1840 m, Schneeweiß, G. & Jang, C.-G. 7764 - https://www.ncbi.nlm.nih.gov/nuccore/AY209285, 2002-06-15" (WU 0032431 - https://wu.jacq.org/WU0032431 - https://www.jacq.org/detail.php?ID=39026 - https://www.gbif.org/occurrence/1830976704; rev. Ó. Sánchez Pedraja Nov 2011 (Schneeweiss & al. 2004)); "Phelypaea tournefortii Desf. / Erzurum: Karaçoban, Kırmızı Tuzla surroundings, steppe, 03 vi 2006, 39° 17' 377'' N 42° 06' 329'' E, 1592 m, LFM 770, Ir.-Tur., Hc. On Centaurea schischkinii [erroneous host-plant?]" (Behçet & al. 2009: 147, sub P. tournefortii); "Erzurum" (photo 69109 by Hayati Keser in www.treknature.com, accessed Mar 2012); "Cat. 2, 1e prijs, Rinus Bode, Phelypaea tournefortii, Kaskargebergte [Pontisch Gebergte / Kaçkar Dağları / Pontic Mountains / Kuzey Anadolu Dağları / North Anatolian Mountains], N-O Turkije" (photo 01 by NRV - Nederlandse Rotsplanten Vereniging - fotowedstrijd van 2012; accessed Feb 2013); "Fotografováno dne 8.6.2013 (Turecko, oblast Aşkale [Erzurum Province], na úbočí Kop Geçidi [Bayburt Province])" (https://botany.cz/cs/phelypaea-tournefortii/ by Pavla Dvorská-Hochmanová, sub P. tournefortii, Autor Eva Rencová, 30.6.2013 in Botany.cz; 2007-2014; accessed Jan 2015); "Narman" (https://gramhir.com/media/2379109340939464334, https://gramhir.com/media/2327631659157093838 and https://gramhir.com/media/2528694368643309541 by @doganinsirdaslari, sub P. coccinea; accessed et rev. Ó. Sánchez Pedraja Aug 2021, sub P. tournefortii, apparently on Tanacetum sp.).

Kars Province / Kars ili: "im Distr. Kaghyzman [Kagizman / Kağızman (Armenian Highlands)] am Sattel Kara dagh (S. Michailowsky)" (Beck 1930: 44, sub P. tournefortii); 1. "Davis [P.H.] & Hedge [J.C.] (D, 29480). / Turkey. Prov. Kars: (Yaqmurlu Daq between / Karaurgan & Sarıkamış, 2200 m. / S. igneous / slopes, parasitic on Tanacetum 29485). / Flowers scarlet with 2 black bosses at / base of lower lip!! Placentation / parietal! 13.6.1957". – 2. "Herb. Musei Hist. Natur. Vindob. / Anoplanthus tournefortii (Desf.) G. Don / 1966 rev. A. Gilli" (https://data.rbge.org.uk/herb/E01283811 - E 01283811.zip - https://www.gbif.org/occurrence/4420516371; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii); 1. "Davis [P.H.] & Hedge [J.C.] (D, 29480). / Turkey. Prov. Kars: (Yaqmurlu Daq between / Karaurgan & Sarıkamış, 2200 m. / S. igneous / slopes, parasitic on Tanacetum 29485). / Flowers scarlet with 2 black bosses at / base of lower lip!! Placentation / parietal! 13.6.1957". – 2. "Phelypaea tournefortii in Kew" (http://specimens.kew.org/herbarium/K004381259 - K 004381259.jpg - https://records.data.kew.org/occurrences/8d6ca828-13ac-4ebe-93d3-0c3862e23861 - https://www.gbif.org/occurrence/5060749994; rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii); 1. "... / Davis [P.H.] & Hedge [J.C.] (D. 30822). / Phelipaea tournefortii Desf. / cited Fl. Turkey / Turkey. Prov. Kars: Yaqmurlu Daq between / Sarıkamış & Karaurgan, 2200 m. Igneous / slopes. On Tanacetum (D. 30698). / 7.6.1957" (http://specimens.kew.org/herbarium/K004381256 - https://records.data.kew.org/occurrences/de7b6340-2e85-40bc-8df6-f3ab7c99af59 - K 004381256.jpg - https://www.gbif.org/occurrence/5060747981; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii [poorly preserved material, with hairy stem and calyx]); "Phelipaea tournefortii Desf., Prov. Kars: Yaqmurlu Daq between Karaurgan & Sarıkamış, 2200 m. 13.06.1957, Davis, P.H., Hedge, J.C. 29480" (http://specimens.kew.org/herbarium/27974.000 - https://www.gbif.org/occurrence/912025203 [n.v.], https://data.rbge.org.uk/herb/E00227405 - https://www.gbif.org/occurrence/1228038038 [n.v., sub Anoplanthus coccineus] et https://data.rbge.org.uk/herb/E00222695 - https://www.gbif.org/occurrence/1228044359 [n.v. (but see K 004381259), sub Anoplanthus coccineus]); "A9 Kars: Yağmurlu Da. [Mount Yagmurlu], 2200 m, D. 30822!" (Gilli 1982: 2, sub P. tournefortii); "Anoplon tournefortii (Desf.) G. Don / Kağızman, above Paslı köyü (Kars) / on Achillea 19971) / 2.VI.1971 Alt: 1800 m / Leg.: T. Baytop ... / No.: 19772" https://data.rbge.org.uk/herb/E01283805 - https://data.rbge.org.uk/search/herbarium/E01283805.zip - https://www.gbif.org/occurrence/4420517347; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii); "Kars [40°14'05.9"N 43°13'24.4"E 40.234980, 43.223430, near Akyayla (Kağızman District [formerly Kaghzvan])], May 2025" (https://www.inaturalist.org/observations/285038006 - https://www.gbif.org/es/occurrence/5177024096 by https://www.inaturalist.org/people/frankgaude [Frank Gaude], sub P. coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2026, sub P. tournefortii, plants with scales only at the base).

Unknown location: 1. "Herbarium Kewense. / Ex Legato Alexander Prior. / Mense Aprili, 1903". – 2. "Anoplanthus Tournefortii Wal. / Kohrab Turkey". – 3. "Orobanche coccinea subsp. coccinea in Kew". – 4. "Diphelypaea coccinea (M.Bieb.) Nicolson in Catalogue of Life" (http://specimens.kew.org/Herbarium/K004381251 - K004381251 - https://records.data.kew.org/occurrences/578c55d3-ef90-4129-9919-b24afcc60c17; rev. Ó. Sánchez Pedraja Aug 2026, sub P. tournefortii).

inicio / top of page

References.

Aznavour, G.V. 1917. Étude sur l'Herbier Artistique Tchitouny (Plantes d'Orient). Mag. Bot. Lap. 16(1-12): 1-37. URL: https://www.zobodat.at/pdf/Ungarische-Botanische-Blaetter_16_0001-0037.pdf.

Bakis, Y., Babac, M.T., & Uslu, E. 2011. Updates and improvements of Turkish Plants Data Service (TÜBİVES). In Health Informatics and Bioinformatics (HIBIT), 2011 6th International Symposium on (pp. 136-140). IEEE. TÜBİVES. Türkiye Bitkileri Veri Servisi [Turkish Plants Data Service]. URL: http://194.27.225.161/yasin/tubives/index.php - http://194.27.225.161/yasin/tubives/index.php?sayfa=1&tax_id=7340 (accessed Sep 2017).

Beck, G. 1930 [30 Sep 1930]. IV. 261. Orobanchaceae Lindl. In: Engler, A. (ed.). Das Pflanzenreich. Regni Vegetabili Conspectus. Wilhelm Engelmann. Leipzig, pp. 1-348. URL: https://www.biodiversitylibrary.org/item/71652.

Behçet, B., Özgökçe, F., Ünal, M. & ve Karabacak, Ö. 2009. The flora of Kırmızı Tuzla (Karaçoban, Erzurum/Turkey) and Bahçe Tuzlası (Malazgirt, Mus/Turkey) and their environment. Biological Diversity and Conservation 2(3): 122-155. URL: https://www.researchgate.net/publication/274721110 - https://www.academia.edu/30756558.

Bizim Bitkiler. 2021. Version 3.1. URL: http://www.bizimbitkiler.org.tr - http://www.bizimbitkiler.org.tr/v3/demo/details.php?id=9705 (accessed Sep 2021).

Boissier, P.É. 1879 [Apr-May 1879]. Flora orientalis [...] Volumen quartum. Fasciculus secundus. Genevae et Basileae. URL: https://bibdigital.rjb.csic.es/idurl/1/10750. [Ord. LXXXIX. Orobanchaceae: 492-518].

Boissier, P.É. & Buhse, F.A. 1860. Aufzaehlung Der Auf Einer Reise Durch Transkaukasien Und Persien Gesammelten Pflanzen. Nouv. Mém. Soc. Imp. Naturalistes Moscou 12. Moscou. Imp. W. Gautier. [Orobanchaceae (*): 168-169 (*) A. Cl. Reutero determinatae]. URL: https://www.biodiversitylibrary.org/item/43404#page/251/mode/1up.

Botany.cz. [Ladislav Hoskovec (Gaston) (Editor in Chief)]. URL: www.botany.cz - https://botany.cz/cs/phelypaea-tournefortii/ (accessed Jan 2015).

Carlón, L., Gómez Casares, G., Laínz, M., Moreno Moral, G., Sánchez Pedraja, Ó. & Schneeweiss, G.M. 2005 [Jun 2005]. Más, a propósito de algunas Orobanche L. y Phelipanche Pomel (Orobanchaceae) del oeste del Paleártico [More on some Orobanche L. and Phelipanche Pomel (Orobanchaceae) from the western Palearctic]. Documentos Jard. Bot. Atlántico (Gijón) 3: 1-71 (in Spanish, plus an Appendix in English).

Carlón, L., Gómez Casares, G., Laínz, M., Moreno Moral, G., Sánchez Pedraja, Ó. & Schneeweiss, G.M. 2008 [17 Jul 2008]. Más, a propósito de algunas Phelipanche Pomel, Boulardia F. W. Schultz y Orobanche L. (Orobanchaceae) del oeste del Paleártico [More on some Phelipanche Pomel, Boulardia F.W. Schultz and Orobanche L. (Orobanchaceae) from the western Palearctic.]. Documentos Jard. Bot. Atlántico (Gijón) 6: 1-128 (in Spanish, plus an enlarged abstract in English).

Cullen, J. 2010 [26 Jan 2010 online]. Phelypaea boissieri: Orobanchaceae. Curtis's Botanical Magazine 26(4): 379-388. DOI: https://doi.org/10.1111/j.1467-8748.2009.01673.x.

Demirkuş, N. [latest update: 21.07.2007]. Phelypaea coccinea and Phelypaea tournefortii. In: The Virtual Herbarium of Van Province. Yüzüncü Yıl University Faculty of Education. Division of Biology Education. Türkçe Van Gölü Havzası Florası (The Flora of Van Lake Basin). Yüzüncü Yıl University Research Council and The Environment Management of Van Province. URL: http://www.vanherbaryum.yyu.edu.tr/ (accessed Jul 2010).

Desfontaines, R.L. 1807-1808. Choix de plantes du Corollaire de Tournefort. Ann. Mus. Par. 10: 218-229, 294-306, 427-433 (1807); 11: 51-57, 136-143, 160-169, 273-282, 376-383, 438-446 (1808); 12: 52-60, 111-118 (1808). URL: http://www.biodiversitylibrary.org/item/23268.

Desfontaines, R.L. 1808. Choix des plantes du Corollaire des Instituts de Tournefort, publiées d'après son herbier, et gravées sur les dessins originaux d'Aubriet. Paris, Levrault. [Phelypaea tournefortii Desf. (1808: 16, t. 10)]. URL: https://archive.org/details/bub_gb_2kUAAAAAQAAJ/page/n21 - https://archive.org/details/bub_gb_60_Uvs9Lv_0C.

Domina, G. & Raab-Straube, E. 2010. Phelypaea tournefortii. Orobanchaceae. In: Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity (accessed Jul 2010).

Don, G. 1837-1838. A general history of the dichlamydeous plants [...] 4 (Corolliflorae). London. [4(1): p. [iii]-viii, [I]-?. 1837; 4(2): p. [i]-iv, ?-908. 8 Mar-8 Apr 1838 (TL-2 Online; IPNI). Order CLXXI. Orobáncheae: 627-636]. DOI: https://doi.org/10.5962/bhl.title.502.

Ekrtová, E. & Ekrt, L. 2012. Křížem krážem arménským Malým Kavkazem II. Nelesní formace - od polopouští po úpatí vysokohorských svahů (Criss-crossing the Armenian Lesser Caucasus II. Non-forest Formations – from Semi-desert to the Foothills of the Great Mountain SlopesCriss-crossing the Armenian Lesser Caucasus II. Non-forest Formations – from Semi-desert to the Foothills of the Great Mo). Živa 2/2012: 69-72. URL: http://botanika.prf.jcu.cz/systematics/publikace/2012_Ziva_Armenie2.pdf.

Fischer, F.E.L. von. 1852. Notice sur les Anoplanthus de l'ancien monde. Bull. Soc. Nat. Mosc. 25(1-2): 105-107. URL: https://www.biodiversitylibrary.org/item/107022.

Gagnidze, R., 2005. Vascular plants of Georgia: A nomenclatural checklist. Tbilisi, "Universale", pp. 1-247.

Gardner, C. & Gardner, B. 2014. Flora of the Silk Road (An Illustrated Guide). I.B. Tauris. London. ISBN: 9781780769417.

GBIF (The Global Biodiversity Information Facility). GBIF Online Resource Centre. Search. Orobanchaceae. URL: https://www.gbif.org/ - https://www.gbif.org/es/species/8348932 (accessed Apr 2018).

Gilli, A. 1971. Die Orobanchaceen der Türkei. Feddes Repert. 82(6): 381-406. DOI: http://dx.doi.org/10.1002/fedr.19710820602.

Gilli, A. 1982. Orobanchaceae (90). In: Davis, P.H. (ed.). Flora of Turkey 7: 1-22. Edinburgh.

Greuter, W., Burdet, H.M. & Long, G. 1989. Med-Checklist.. 4: 265-266 [Orobanchaceae]. Genève, Berlin.

iNaturalist. URL: https://www.inaturalist.org/ - https://www.inaturalist.org/taxa/ - https://www.inaturalist.org/observations?taxon_id=128605 - https://www.inaturalist.org/taxa/129838 - https://www.inaturalist.org/taxa/128605 - https://www.inaturalist.org/taxa/128603 (accessed Feb 2020).

Index Synonymique de la Flore de France. Kerguélen, M. 1999. [J.-P. Lonchamp, Oct. 1999, Electronic version]. 1998-2002 - INRA-MNHN. URL: http://www.dijon.inra.fr/flore-france/index.htm (accessed Sep 2007).

Karjagin, I.I. (ed.). 1957. Flora Azerbajdžana [Azerbajcan Florasy] 7 (Pyrolaceae-Plantaginaceae). Baku: Akademija Nauk Azerbajdzanskoj SSR, Institut botaniki imeni V.L. Komarova.

Lambert, A.B. 1811. VII. Some Account of the Herbarium of Profesor Pallas. Trans. Linn. Soc. London 10: 256-265. [Phelipaea foliata: 260-262]. URL: https://www.biodiversitylibrary.org/item/13720.

Leonidovich, M.S. [Леонідович, М.С.]. 2016. ПАЛІНОМОРФОЛОГІЧНІ ОСОБЛИВОСТІ ПРЕДСТАВНИКІВ ПОРЯДКУ Lamiales L.: ФІЛОГЕНЕТИЧНЕ ЗНАЧЕННЯ ТА НАПРЯМ КИ ЕВОЛЮЦІЇ. 03.00.05 - ботаніка. Дисертація на здобуття наукового ступеня доктора біологічних наук. Київ [Kiev]. УДК 581.33:582.916.21:575.85/.86.

Linnaeus, C. 1758. Opera varia in quibus continentur Fundamenta botanica, Sponsalia plantarum, et Systema naturae: in quo proponuntur naturae regna tria secundum classes, ordines, genera & species. Lucae: Ex typographia Juntiniana, 1758. [Op. Var.]. DOI: https://doi.org/10.5962/bhl.title.154190. URL: https://www.biodiversitylibrary.org/bibliography/154190. (TL-2: 101 n. 4798).

MNHN (Muséum National d'Histoire Naturelle), Paris (France). Vascular plants (P). Orobanchaceae. URL: https://science.mnhn.fr/all/search - https://science.mnhn.fr/institution/mnhn/collection/p/item/search - http://coldb.mnhn.fr/catalognumber/mnhn/p/p02970941 [permalink]... - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970941 [browser link]... - http://mediaphoto.mnhn.fr/media/1441352983821Tbv218qAE1INSMiF [image link]... (accessed Nov 2011).

Naturalis - Nationaal Herbarium Nederlands - Botany Section (AMD - L - WAG). BioPortal. Naturalis Biodiversity Center (NL). URL: http://data.biodiversitydata.nl/naturalis/specimen/L.2821536 [permalink]... - http://medialib.naturalis.nl/file/id/L.2821536/format/large [image link]... (accessed Oct 2017).

Nicolson, D.H. 1975 [Nov 1975]. Diphelypaea (Orobanchaceae), nom. nov. and other cauterizations on a nomenclatural hydr. Taxon 24(5-6): 651-657. DOI: https://doi.org/10.2307/1220740.

Novopokrovskij, I.V. & Tzvelev, N.N. 1958. Orobanchaceae. In: Shishkin, B.K. (ed.). Flora Unionis Republicarum Socialisticarum Sovieticarum (Flora URSS), vol. 23. Institutum Botanicum nomine V. L. Komarovii Academiae Scientiarum URSS, Mosqua and Leningrad, pp. 685–687. (Novopokrovskii, I.V. & Tzvelev [Cvelev], N.N. 1958. Orobanchaceae Vent. In: Komarov, V. L., (Ch. Ed.). 1934-1964. Flora SSSR 23: 17-111. Edit. Akad. Nauk SSSR. Moskva-Leningrad. [Pandit, V. (Translator & Editor). 1999. Amerind Publishing Co. Pvt. Ltd., New Delhi. Translated and published under an agreement with the Smithsonian Institution Libraries, Washington, DC, by Amerind Publishing Co., 66 Janpath, New Delhi 110 001. 2000] et [Panigrahi, G. (Bot. Ed.). 2000. Bishen Singh Mahendra Pal Singh]).

Piwowarczyk, R., Sánchez Pedraja, Ó., Moreno Moral, G., Fayvush, G., Zakaryan, N., Kartashyan, N. & Aleksanyan, A. 2019 [3 Jan 2019]. Holoparasitic Orobanchaceae (Cistanche, Diphelypaea, Orobanche, Phelipanche) in Armenia: distribution, habitats, host range and taxonomic problems. Phytotaxa 386(1): 001-106. DOI: https://doi.org/10.11646/phytotaxa.386.1.1.

Piwowarczyk, R., Sánchez Pedraja, Ó., Khutsishvili, M. & Kharazishvili, D. 2023 [24 Jul 2023]. Holoparasitic Orobanchaceae in Georgia (Caucasus): taxonomic revision, diversity, distribution, habitats and host range. Phytotaxa 604(1): 1-103. DOI: https://doi.org/10.11646/phytotaxa.604.1.1. URL: https://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.604.1.1 - https://www.researchgate.net/publication/372570616.

Plantarium. Открытый атлас сосудистых растений России и сопредельных стран [Open Atlas of Vascular Plants of Russia and Adjacent Countries]. Определитель растений on-line [To Plants of the On-line]. Поиск таксонов [Search taxa]. Scrophulariaceae. Cistanche. Diphelypaea. Orobanche. URL: http://www.plantarium.ru/ (accessed Jan 2015).

Reichenbach, H.G.L. 1823-1832. Iconographia botanica seu plantae criticae [...] Septima centuria tabularum: 29-50, t. 651-700 [DCLI-DCC] (1829). Lipsiae: Apud Fridericum Hofmeister. URL: https://bibdigital.rjb.csic.es/idurl/1/10431. (TL-2: 670 n. 8876; Kergélen, 1987: 136, "1931", Greuter, 1989: 38, "1931").

Reuter, G. 1847 [25 Nov 1847]. Orobanchaceae. In: De Candolle, A. (ed.). Prodromus Systematis naturalis regni vegetabilis... 11: 1-45. Addenda et Corrigenda: 717-720. Parisiis: Sumptibus Victoris Masson. URL: https://www.biodiversitylibrary.org/page/160970. (TL-2: 446 n. 999).

Reveal, J.L. 1991. Two previously unnoticed sources of generic names published by John Hill in 1753 and 1754-1755. Bull. Mus. Natl. Hist. Nat.ser. 4, sect. B Adansonia 13: 197-239. [Phelypaea: 228].

Rix, E.M. & Webb, D.A. 1972. Phelypaea L. In: Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (eds.). Flora Europaea 3: 294.

Schneeweiss, G.M., Palomeque, T., Colwell, A.E. & Weiss-Schneeweiss, H. 2004 [Mar 2004]. Chromosome numbers and karyotype evolution in holoparasitic Orobanche (Orobanchaceae) and related genera. American Journal of Botany 91: 439-448. [Abstract / Download. Supplemental Data: Appendix 1 - New chromosome counts for species of Cistanche, Diphelypaea, and Orobanche with collection details. Appendix 2 - Chromosome numbers of Conopholis, Cistanche, Diphelypaea, and Orobanche] (accessed Sep 2007).

Schneeweiss, G.M., Colwell, A.E., Park, J.M., Jang, C.G. & Stuessy, T.F. 2004. Phylogeny of holoparasitic Orobanche (Orobanchaceae) inferred from nuclear ITS sequences. Mol. Phylogenetics Evol. 30: 465-478. DOI: https://doi.org/10.1016/S1055-7903(03)00210-0. [Abstract / Download].

Schneider, A.C. 2016 [9 Dec 2016]. Resurrection of the genus Aphyllon for New World broomrapes (Orobanche s.l., Orobanchaceae). PhytoKeys 75: 107-118. DOI: https://doi.org/10.3897/phytokeys.75.10473.

Seregin, A.P. (ed.). 2017. Specimen MW0723306... from the collection "Moscow University Herbarium" // Depository of Live Systems (branch "Plants"): Electronic resource. – Moscow State University, Moscow. URL: https://plant.depo.msu.ru/open/public/item/MW0723306 [permalink]... - https://plant.depo.msu.ru/open/public/item/MW0723306/img/0.jpg?original=253c396e-7bd9-42c3-99d4-0092276e4174 [image link]... - https://plant.depo.msu.ru/open/public/item/MW0723306/img/0.jpg?original [former image link]... - https://plant.depo.msu.ru/open/public/item/MW0723306/img/0.jpg?gbif [image link, accessed via GBIF.org]... (accessed Apr 2018).

SIUC (Parasitic Plant Connection). SIUC / College of Science / Parasitic Plant Connection / Scrophulariaceae. URL: http://www.parasiticplants.siu.edu/Scrophulariaceae/NoPhoto.Scrophs.html.  Last updated: 07-Dec-15 / dln (accessed Sep 2017).

Sorger, F. 1994. Blumen der Türkei. Stapfia 34: 103-271. URL: https://archive.org/details/stapfia-34-103-271 - https://www.zobodat.at/pdf/STAPFIA_0034_0103-0271.pdf.

Sprengel, C.P.J. 1824-1828 [Jan-May 1825]. Systema vegetabilium [Caroli Linnaei ... ]. (Editio decima sexta) 2. Gottingae. [2205. Orobanche: 815. 2206. Phelipaea: 818 ... 2211. Aeginetia: 819. Syst. Veg. (ed. 16) 2 (1825)]. DOI: https://doi.org/10.5962/bhl.title.822. (IPNI and TL-2).

Stapf, O. 1915. The Genus Phelipaea. Bull. Misc. Inform. Kew [Bulletin of Miscellaneous Information (Royal Gardens, Kew)] 1915, 6: 285-295. URL: https://www.biodiversitylibrary.org/item/42572.

Steudel, E.G. 1840-1841 [Apr 1841]. Nomenclator botanicus seu: Synonymia plantarum universalis, enumerans ordine alphabetico nomina atque synonyma tum generica tum specifica et a Linnaeo et a recentioribus de re botanica scriptoribus plantis phanerogamis imposita. Stuttgartiae et Tubingae: Typis et sumptibus J. G. Cottae. DOI: https://doi.org/10.5962/bhl.title.655. URL: https://bibdigital.rjb.csic.es/idurl/1/10860 [Nomencl. Bot. ed. 2.2]. [Orobanche Lin. Spr. 2205.: 230-232]. (TL-2: 909 n. 13053).

Tarkhnishvili, D. & Chaladze, G. (eds.). 2013. Georgian biodiversity database. URL: http://www.biodiversity-georgia.net/index.php?taxon=Diphelypaea%20tournefortii.

Teryokhin, E.S., Shibakina, G.V., Serafimovich, N.B. & Kravtsova, T.I. 1993. Opredelitelj Sarasichovich Florii SSSR (Determinator of Broomrapes of the USSR Flora). Leningrad.

The Caucasus. Coordination and Development of Plant List Assessments for the Caucasian Biodiversity Hotspot. IUCN (The World Conservation Union), Missouri Botanical Garden] & al. Critical Ecosystem Parnership Fund (CEPF) (accessed Feb 2009).

Tournefort, J.P. de. 1700. Institutiones rei herbariæ. Parisiis, E Typographia regia, 1700-[1703]. [Editio altera: 1700, 3 vols.]. DOI: https://doi.org/10.5962/bhl.title.713 [1: Genus IX Orobanche. 175-176. 3: Phelypaea t. 479].

Tournefort, J.P. de. 1703. Corollarium Institutionum rei herbariæ, Plantae 1356. munificentiâ Ludovici Magni in Orientalibus Regionibus observata recensentur, & ad genera sua revocantur. Parisiis: E Typographia Regia, pp. [i-viii], [1]-58. [N.B. It is a supplement, published in 1703, to the Instututiones Rei Herbariæ]. [Orobanche: 10; Phelypaea: 47]. DOI: https://doi.org/10.5962/bhl.title.713.

Tzvelev, N.N. 1981 [2001]. Orobanchaceae Vent. In: Fedorov, An.A. (ed.). Flora of Russia: The European Part and Bordering Regions 5: 434-461. Nauka Publishers. Leningrad [A.A. Balkema. India]. [Flora Evropeiskoi Chasti SSSR 5].

Utah State University. 2019. Intermountain Herbarium (Utah State University). Occurrence dataset https://doi.org/10.15468/t43wjj accessed via GBIF.org on 2020-02-05. https://www.gbif.org/occurrence/1986951694.

Wallroth, K.F.W. 1822. Schedulae criticae de Plantis Florae Halensis selectis. Halae (Halle an der Saale): sumtibus Car. Aug. Kümmelii. [XXIX. Orobanche Linn.: 304-315]. DOI: http://dx.doi.org/10.3931/e-rara-17327 (TL-2 16.595).

Weiss-Schneeweiss, H., Greilhuber, J. & Schneeweiss, G.M. 2006. Genome size evolution in holoparasitic Orobanche (Orobanchaceae) and related genera. Am. J. Bot. 93: 148-156. [Abstract / Download]. DOI: https://doi.org/10.3732/ajb.93.1.148. URL: http://www.amjbot.org/content/93/1/148.full.pdf+html.

Willdenow, C.L. 1800. Caroli a Linné species plantarum exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus secundum systema sexuale digestas. Berolini: Impensis G. C. Nauk. Tomus III. Pars I. [Sp. Pl. 3(1)] [1186. Orobanche: 347-354]. URL: https://bibdigital.rjb.csic.es/idurl/1/12398.

Zare, G., Dönmez, A.A. & Dönmez, E.O. 2014 [May 2014]. Pollen morphology and evolution in the genus Orobanche L. s.l. and its allied genera (Orobancheae/Orobanchaceae) in Turkey. Plant. Syst. Evol. 300(5): 783-802. DOI: https://doi.org/10.1007/s00606-013-0919-2.

Zare, G. & Dönmez, A.A. 2016 [Jan 2016]. Fruit and seed morphology of the tribe Orobancheae (Orobanchaceae) genera in Turkey and its taxonomic significance. Nordic J. Bot. 34(2): 178-190. DOI: https://doi.org/10.1111/njb.00945.

inicio / top of page

Home page (Página principal) | Index of Orobanchaceae | Contact us

© 2005 GBC (Grupo botánico cantábrico)