Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us ISSN: 2386-9666 - Last Updated: 11 September 2023 |
Accepted Name: Phelipanche gratiosa (Webb) Carlón, G. Gómez, M. Laínz, Moreno Mor., Ó. Sánchez & Schneew. in Documentos Jard. Bot. Atlántico (Gijón) 6: 66[-67], 63 [key], f. 1 [phylogenetic relationships] (2008 [17 Jul 2008]) Basionymum / Basionym: ≡ Phelypaea gratiosa Webb in Webb & Berthel., Phytogr. Can. 3: 153[-154] (1845 [Dec 1845]) ["Phelipaea"] (Reuter 1847: 6, sub "P. graciosa") Synonyms: Homotypic synonyms: ≡ Orobanche gratiosa (Webb) Linding. in Abh. Gebiet der Auslandskunde 21: 248 (1926) (view IPNI entry, accessed Feb 2006) ≡ Orobanche gratiosa (Webb) Beck in Engl., Pflanzenr. 96: 112 (1930) ["? O. graciosa (Webb et Berth.) G. Beck"], comb. superfl. Misapplied names – Orobanche mutelii auct. canar. pl., non Orobanche mutelii F.W. Schultz in Mutel, Fl. Franç. 2: 353 (1835) (Marrero & al. 1995: 103) – Phelypaea ramosa auct. canar. pl., non Orobanche ramosa L., Sp. Pl.: 633 (1753) (Plantae Canarienses - hb. C.J. Pitard, 1905: P 02970683 - Pitard n. 312, 1905; P 02970684 - Pitard n. 1789, 1905) |
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Ind. loc.: Spain / España, Canary Islands: "Hab. Occurrit haec species in Lancerottá et in insulà arenosa cui nomen Graciosa [Lanzarote Island and Graciosa Island]. / Stir pern nutricem cujus parasita non licuit nobis secernere." (Webb 1845: 154). Lectotypus / Lectotype: "Herbariun / Nº 141629 Webbianum / Herbarium / Universitatis Florentinae / FI 000239 / Typus / Fototeca / N.º 2258 / Data 3-5-07 [2007]) // "Orobanche / in Lancerotta" // "Orobanche gratiosa Nob." // "Ph. Salviae aff." // "Orobanche gratiosa (Webb & Berthelot) Lindinger / Typus-lecto / Determinavit A. Santos" (FI 000239; rev. Ó. Sánchez Pedraja Mar 2012, lectotype of P. gratiosa (Webb 1845: 154) [the sheet contains two specimens, the left-hand of the pair is a suitable lectotype for this species. Moreover, this sheet bears various labels]). "Orobanche è Canariis insulis / Phelipaea gratiosa Webb et Berthel. Phytogr. canar. / sect. 3, p. 153 (82 ... publ. le 13 Dec. 1845). — Reut. in Decand. prodr / XI (1847) p-6, nº. 7 / Webb & Berthelot / dederunt Junio 1834" (K 000425197 - hb. J. Gay; rev. Ó. Sánchez Pedraja Mar 2012, isolectotypus?). Designated by: A. Santos [2 Dec 1995]. Host: habitually (in at least 90 percent of cases) on Launaea arborescens (Batt.) Murb. [≡ Zollikoferia arborescens Batt.; – Prenanthes spinosa auct., non Forssk. (Bourgeau 1846, in sched.)], Compositae. More rarely on other Compositae species [Reichardia tingitana (L.) Roth; Urospermum picroides ( L. ) F.W.Schmidt]. Coincidentally on other families, e. g., on Zygophyllaceae (Fagonia cretica L.) and on Graminae (Arundo donax L.) (Annotated Checklist of Host Plants of Orobanchaceae, accessed Apr 2010). Comments:
Chromosomatic number: n = 12, 2n = 24 (Schneeweiss & al., 2004, sub "cf. gratiosa G. Beck"). Ic.: a, inflorescences (on Launaea arborescens (Batt.) Murb.; Lanzarote, Mala; terr. class.); b, flowers (ib.); c, seeds (ib.); d, sheet (ib.); e, flower sketches: P. nana vs. mutelii vs. aedoi vs. gratiosa; f, Canariis insulis, leg. Webb & Berthelot, 1834 (K 000425197 - hb. J. Gay, isolectotypus?); g, Fuerteventura, Puerto de Cabras (P 02970683 - Pitard n. 312, 1905); h, Lanzarote, Haria (P 02970684 - Pitard n. 1789, 1905). Distribution: Canary Islands distribution. The Phelipanche gratiosa is endemic to the Canary Islands. |
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Specimens examined and literature records. Spain / España / Espagne / Spanien Canarias, Islas / Canary Islands / Kanarischen Inseln / Îles Canaries (Webb 1845: 154, sub Phelypaea gratiosa; Lindinger 1926: 248, sub O. gratiosa) Fuerteventura: 1. "Orobanche / da Prenanthes spinosa [Launaea arborescens (Batt.) Murb.] / Partie Sud de Handia / Fuerteventure 26 Febrero 1846". – 2. "E. Bourgeau, Plantas Canarienses nº. 935 / Orobanche gratiosa Webb (Phyt. ...) / Fuerteventura — in arenis Handia [Jandía, (pronounced "handia")] die 26 Februarii 1846" (FI - W 141630, the two perfect specimens on the right-hand of the sheet; rev. L. Carlón & Ó. Sánchez Pedraja 2006-2007); “E. Bourgeau, Plantas Canarienses nº. 935 / Orobanche gratiosa Webb ? / Fuerteventura — In arenis Handia [Jandía, (pronounced "handia")] / die 26 Februarii 1846” (P 02970717; rev. Ó. Sánchez Pedraja Mar 2012); 1. "Plantae Canarienses / 312 Phelippaea ramosa. L. / Fuerteventura: Puerto de Cabras [Puerto del Rosario], in aridis maritimis. / 22 2 1905 C.J. Pitard". – 2. "Herb. Mus. Paris. / Herb. J. Pitard (Canaries), donné par Mme Pitard / Reçu le 16 Mai 1929" (P 02970683 - Pitard n. 312, 1905; rev. Ó. Sánchez Pedraja Mar 2012); 1. "Plantae Canariensis / 312 Phelipaea ramosa. L. / Fuerteventura: Puerto de Cabras, / in aridis maritimis / 22.2. 1905. C.J. Pitard". – 2. "Orobanche ramosa L. / I.K. 1965". – 3. "Orobanche mutelii Schultz / Det. F.M. Muller 1986 Herbarium Vadense (WAG)" (WAG 1141540; rev. Ó. Sánchez Pedraja Jun 2017); Puerto del Rosario-La Oliva, V. Montelongo, 18 Feb.1984 (LPA 013815, sub O. cf. ramosa, rev. sub O. mutelii F.W. Schultz, A. Pujadas II-1997; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); "Jopo de la Graciosa (Phelipanche gratiosa) ... proximidades del Pico de la Zarza [Pico de Jandía]" (photo 2015/09/Jopo-azul-2-2-.jpg in Fuerteventura en imágenes, accessed Jun 2017); Spain - Fuerteventura - Corralejo (s), 2023-01-25" (https://observation.org/observation/262649308 - https://www.gbif.org/occurrence/4057990064 by Edo Kreuzen & Hilde Ham, sub O. minor Sm., in Observation.org; accessed et rev. Ó. Sánchez Pedraja Sep 2023, sub P. gratiosa); "Spain - Fuerteventura - Jable de Corralejo, 2023-01-25" (https://observation.org/observation/262639476 - https://www.gbif.org/occurrence/4056904290 by Jacques van Krieken, sub O. minor Sm., in Observation.org; accessed et rev. Ó. Sánchez Pedraja Sep 2023, sub P. gratiosa). Graciosa (La): in insulà arenosa cui nomen Graciosa (Webb 1845: 154); Teguise, "entre el Mojón y nuestra Señora de las Agujas", A. Marrero, 26 Mar 1988 (LPA 016963, sub Orobanche, rev. sub Orobanche mutelii F.W. Schultz, A. Pujadas Feb 1997; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007). Gran Canaria: "Orobanche / da Prenanthes spinosa [Launaea arborescens (Batt.) Murb.] / Sur les collines du Fort Royal / de Gde. Canaria 14 Mars 1846", [E. Bourgeau N.º 1045] (FI - W 141630, the problematic specimen [cf.] on the left-hand of the sheet; rev. L. Carlón & Ó. Sánchez Pedraja 2006-2007); Laderas del Jardín Botánico "Viera y Clavijo", A. Marrero & M. González-Martín, 7 Mar. 1995 (LPA 018250, COA et MA, sub Orobanche mutelii F.W. Schultz; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); slope of the Jardín Canario, 275 m, Las Palmas de Gran Canaria, A. Marrero, 4 Mar 1997 (LPA 018283 et COA, sub Orobanche mutelii F.W. Schultz; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); San Nicolás de Tolentino, G. Gómez Casares, apparently parasitic on Launaea arborescens (Batt.) Murb. (Compositae) in volcanic soils, 28 Feb 2006 (obs.); "Todas las fotos han sido tomadas en una población de Gran Canaria situada entre el barranco de Venegueras [Mogán] y el de Tasarte [San Nicolás de Tolentino] a unos 800 m.s.n.m." (photos of Phelipanche cf. gratiosa [imgp2662!, imgp2661!, imgp2647!, imgp2671!] by Juanillo in Plantasdemitierra.blogspot.com, accessed Oct 2010). Lanzarote: "Herbariun / Nº 141629 Webbianum / Herbarium / Universitatis Florentinae / FI 000239 / Typus / Fototeca / N.º 2258 / Data 3-5-07 [2007]) // "Orobanche / in Lancerotta" // "Orobanche gratiosa Nob." // "Ph. Salviae aff." // "Orobanche gratiosa (Webb & Berthelot) Lindinger / Typus-lecto / Determinavit A. Santos" (FI 000239; rev. Ó. Sánchez Pedraja Mar 2012, lectotype of P. gratiosa (Webb 1845: 154) [the sheet contains two specimens, the left-hand of the pair is a suitable lectotype for this species. Moreover, this sheet bears various labels]); "Orobanche è Canariis insulis / Phelipaea gratiosa Webb et Berthel. Phytogr. canar. / sect. 3, p. 153 (82 ... publ. le 13 Dec. 1845). — Reut. in Decand. prodr / XI (1847) p-6, nº. 7 / Webb & Berthelot / dederunt Junio 1834" (K 000425197 - hb. J. Gay; rev. Ó. Sánchez Pedraja Mar 2012, isolectotypus?); Occurrit haec species in Lancerottá" (Webb, 1845: 154); 1. "Flore des Canaries / 1789 Phelipaea ramosa. Mey. / Lanzarote: Haria, in aridis / III 1905. C.J. Pitard". – 2. "Herb. Mus. Paris. / Herb. J. Pitard (Canaries), donné par Mme Pitard / Reçu le 16 Mai 1929" (P 02970684 - Pitard n. 1789, 1905; rev. Ó. Sánchez Pedraja Mar 2012); "Lanzarote (WB 50a)" (Lindinger, 1926: 248, sub O. gratiosa); "Herbariu Groninganum D 5349 / Orobanche spec. [sub O. mutelii] / Canarische eilanden / Lanzarote; bergen ten zuiden van Haria / Leg. C.H. Andreas & H.D. Schotsman 24-29.III.1972" (WAG 1141541; rev. Ó. Sánchez Pedraja Jun 2017); 1. "Herbariu Groninganum D 5349 / Orobanche cf. cernua / Det. Andreas / Canarische eilanden / Lanzarote; W. zijde Famara complex / ... / Leg. C.H. Andreas & H.D. Schotsman / III-1972". – 2. "Orobanche barbata Poiret? / Det. F.M. Muller, 1986 Herbarium Vadense (WAG)" (https://data.biodiversitydata.nl/naturalis/specimen/WAG.1141402 - WAG 1141402.jpg - https://www.gbif.org/occurrence/2517714053; rev. Ó. Sánchez Pedraja Jun 2017); Valle de San José, Teguise, A. Marrero & A. Carrasco, 10 June 1995 (LPA 018056 et MA 564183, sub O. mutelii F.W. Schultz (Marrero & al., 1995: 103); rev. L. Carlón, G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); Los Valles, Teguise, A. Marrero & A. Carrasco, 10 June 1995 (LPA 018057 et ORT, sub O. mutelii F.W. Schultz (Marrero & al., 1995: 103); rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); Montaña Halcones, Timanfaya, M. González-Martín & A. Marrero, 12 Mar. 1994 (LPA 018058, sub O. mutelii F.W. Schultz (Marrero & al. 1995: 103); rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); west summit of Termesana mountain, Timanfaya, M. González-Martín & A. Marrero, 19 Mar. 1994 (LPA 018059, sub O. mutelii F.W. Schultz (Marrero & al., 1995: 103); rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); El Taro, Timanfaya, A. Marrero & M. González-Martín, 15 May 1994 (LPA 018065, sub Orobanche et sub O. ramosa “? revisar”; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); Montaña Ganada, Haría, A. Marrero, M. González-Martín, A. Perdomo & A. Carrasco, 11 Mar. 1995 (LPA 018203 et MA 564182, sub O. mutelii F.W. Schultz (Marrero & al. 1995: 103); rev. L. Carlón, G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); Jables [sand dunes] de Famara, el Marichuelo, Teguise, A. Marrero, M. González-Martín & A. Perdomo, 12 Mar. 1995 (LPA 018204 et ORT, sub O. mutelii F.W. Schultz (Marrero & al., 1995: 103); rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); Jables de Famara, Teguise, A. Marrero, M. González-Martín & A. Perdomo, 12 Mar. 1995 (LPA 018252, sub Orobanche; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); Famara-Gebirge, bei Guinate, G. Schneeweiss & H. Weiss-Schneeweiss H. 8757, 2-IV-2002 (WU 0027581, WU 0027582, WU 0027583 et WU 0027584 - Phanerogamen); between Playa Blanca and Punta del Águila, 0-30 m, on Launaea arborescens (Batt.) Murb. (Compositae), G. Schneeweiss & H. Weiss-Schneeweiss (WU: S & T 8753) [AY209331, sub "Orobanche cf. graciosa (Webb & Berth.) Winding"]; between the Mojón and Guatiza [Teguise municipality], 28RFT4617, 175 m, "parasita al género Launaea, especialmente a Launaea arborescens", J.A. Reyes Betancort, 24 Mar. 2005 (ORT; rev. G. Gómez Casares & Ó. Sánchez Pedraja 2006-2007); El Bosquecillo, Lanzarote, M.L. Gil González; Teguise municipality, Costa Teguise (near Teguise), 28RFT4809, 10 m, apparently parasitic on Launaea arborescens (Batt.) Murb. in waste ground (sandy and wet soils), G. Moreno Moral, 23 Mar 2006 (obs.); Teguise municipality, 28RFT4709, 10 m; in stony soils of a flowerbed, G. Moreno Moral, 24 Mar. 2006 (obs.); Teguise municipality, between the Montaña de Saga and the Montaña Corona (Costa Teguise, near Teguise village), 28RFT4511, 70 m, apparently parasitic on L. arborescens (Batt.) Murb. in a stream bed with stony soils, G. Moreno Moral MM0020/2006, 25 Mar 2006 (herb. Sánchez Pedraja 12461); Teguise municipality, sand dunes of Famara (near Famara), 28RFT4021, 5 m, apparently parasitic on Launaea arborescens (Batt.) Murb. in a dry stream bed, G. Moreno Moral MM0021/2006, 25 Mar 2006 (herb. Sánchez Pedraja 12462); Teguise municipality, below the Tahíche village (south slope), 28RFT4209, 160 m, apparently parasitic on L. arborescens (Batt.) Murb. in stony soils, G. Moreno Moral, 25 Mar 2006 (obs.); Teguise municipality, near the Cuesta Jai (near Tahíche village), 28RFT3713, 200 m, apparently parasitic on L. arborescens (Batt.) Murb. (cf.) in scrub with sandy soils, G. Moreno Moral, 25 Mar 2006 (obs.); Teguise municipality, Montaña Cobrada (near Guatiza village), 28RFT4918, 70 m; apparently parasitic on L. arborescens (Batt.) Murb., G. Moreno Moral, 26 Mar 2006 (obs.); Teguise municipality, near Guatiza village, 28RFT4818, 80 m; apparently parasitic on L. arborescens (Batt.) Murb. in waste ground (sandy soils), G. Moreno Moral, 26 Mar 2006 (obs.); Teguise municipality, Costa Teguise (near Teguise village), 28RFT4809, 10 m, apparently parasitic on L. arborescens (Batt.) Murb. in sandy and wet soils, G. Moreno Moral MM0049/2006, 30 Mar 2006 (herb. Sánchez Pedraja 12493); Haría municipality, next to the beach of the Caleto Blanco (near Órzola village), 28RFT5231, 20 m, apparently parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0010/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12450); Haría municipality, next to the Barranco de Chafarís (near Tabayesco village), 28RFT4822, 30 m, apparently parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0011/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12451); Haría municipality, next to the Barranco de Chafarís (near Tabayesco village), 28RFT4823, 30 m, apparently parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0012/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12452 - EU581791); Haría municipality, next to the Barranco de Chafarís (near Tabayesco village), 28RFT4823, 30 m, parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0013/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12453); Haría municipality, waste ground with stony soils at the foot of the slope (near Mala village), 28RFT4821, 60 m, apparently parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0016/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12457); Haría municipality, waste ground with stony soils at the foot of the slope (near Mala village), 28RFT4821, 60 m, apparently parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0016A/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12457A); Haría municipality, waste ground with stony soils at the foot of the slope (near Mala village), 28RFT4821, 60 m, parasitic on Reichardia tingitana (L.) Roth, G. Moreno Moral MM0017/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12458); Haría municipality, slope below the reservoir of Mala (near Mala village), 28RFT4821, 120 m, apparently parasitic on L. arborescens (Batt.) Murb. (Compositae), G. Moreno Moral MM0018/2006, 24 Mar 2006 (herb. Sánchez Pedraja 12459); Haría municipality, at the top of the Valle de Malpaso —pr. Haría—, 28RFT4423, 480 m, apparently parasitic on L. arborescens (Batt.) Murb. in stony soils, G. Moreno Moral MM0022/2006, 25 Mar 2006 (herb. Sánchez Pedraja 12463); Haría municipality, 28RFT4423, 440 m, apparently parasitic on L. arborescens (Batt.) Murb. in roadside, G. Moreno Moral, 25 Mar. 2006 (obs.); Haría municipality, at the top of the Valle de Malpaso (near Haría village), 28RFT4423, 480 m, parasitic on (!) Fagonia cretica L. (Zygophyllaceae) in terraced and stony soils, G. Moreno Moral MM0023/2006, 25 Mar 2006 (herb. Sánchez Pedraja 12464); Haría municipality, 28RFT4423, 480 m, parasitic on (!) Urospermum picroides ( L. ) F.W.Schmidt in terraced and stony soils, G. Moreno Moral MM0024/2006, 25 Mar 2006 (herb. Sánchez Pedraja 12465); Haría municipality, at the top of the Valle de Malpaso (near Haría village), 28RFT4423, 480 m, parasitic on (!) Launaea arborescens (Batt.) Murb., in terraced and stony soils, G. Moreno Moral MM0025/2006, 25 Mar 2006 (herb. Sánchez Pedraja 12466); Haría municipality, next to a ravine just south of the Montaña Corona (Costa Teguise, near Teguise village), 28RFT4510, 50 m, apparently parasitic on L. arborescens (Batt.) Murb. in stony soils, G. Moreno Moral, 25 Mar 2006 (obs.); ibid., above Los Valles (next to Km 13 on road to Haría), 28RFT4419, 430 m, apparently parasitic on L. arborescens (Batt.) Murb. (cf.) in stony soils, G. Moreno Moral, 25 Mar 2006 (obs.); Haría municipality, at the top of the ravine of Chafarís (Temisa valley, above Tabayesco village), 28RFT4522, 280 m; apparently parasitic on L. arborescens (Batt.) Murb. in the roadside, G. Moreno Moral, 28 Mar 2006 (obs.); Haría municipality, below the Montaña del Mojón (beside Charco del Palo), 28RFT4918, 60 m; apparently parasitic on L. arborescens (Batt.) Murb. in fixed and inland sand dunes (jables), G. Moreno Moral, 28 Mar 2006 (obs.); Teguise municipality, sand dunes of Famara, 29º06'52''N, 13º33'21''W, C. Aedo, L. Medina & A. Quintanar 12472, 4-IV-2006, parasitic on Launaea en las dunas (MA; det. G. Moreno Moral & Ó. Sánchez Pedraja 2006-2007); Haría municipality, below the Montaña del Mojón (beside Charco del Palo), 28RFT4918, 60 m, apparently parasitic on L. arborescens (Batt.) Murb. in fixed and inland sand dunes (jables), G. Moreno Moral MM0044/2006, 29 Mar 2006 (herb. Sánchez Pedraja 12486); Tinajo municipality, at the foot of the Caldera Blanca (near Casas del Islote), 28RFT2313, 150 m, apparently parasitic on L. arborescens (Batt.) Murb. in waste ground, G. Moreno Moral, 28 Mar. 2006; Tinajo municipality, 28RFT2312, 150 m, apparently parasitic on L. arborescens (Batt.) Murb. in gravel soils, G. Moreno Moral, 28 Mar 2006 (obs.); "Órzola [Haría municipality]. Lanzarote" (photos by M.L. Gil González in Flora Vascular de Canarias, accessed Jul 2010 et Jun 2016); Famara cliffs 18 April 2015 (photo 2015/10/dscn9555.jpg by Brad Scott in A long walk to the cliffs, accessed Jun 2017). |
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