Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

Accepted Name:

Phelipanche coelestis (Reut.) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972)

Basionymum / Basionym:

≡ Phelypaea coelestis Reut. in DC. Prod. 11: 5 (1847) ["Phelipaea"]

Synonyms:

≡ Orobanche coelestis Boiss. & Reut. in Pinard, Pl. Exscc. (1843), nom.illeg. [cf. Reut. (1847: 5, 37); Beck (1930: 93); Novopokrovskii & Tzvelev (1958: 60)]

≡ Phelypaea coelestis Reut. in DC. Prod. 11: 5 (1847) ["Phelipaea"]

≡ Orobanche coelestis Boiss. & Reut. ["in Pin. pl. exs. 1843"] ex Reut. in DC. Prod. 11: 5 [37] (1847), pro syn. [sec. (IK) view IPNI entry, accessed 17 Sept. 2007]

≡ Orobanche coelestis Boiss. & Reut. ex Beck in Biblioth. Bot. 19: 114 (1890) [sec. w3Tropicos, accessed 17 Sept. 2007]

≡ Orobanche coelestis (Reut.) Beck in Biblioth. Bot. 19: 114 (1890) [sec. Greuter & al., 1989: 258; nom. val.(!) sec. ICBN - St. Louis Code, Art. 34]

≡ Kopsia coelestis (Reut.) Bég. & A. Vacc. in Ann. Bot. (Roma) 13: 29 (1914)

≡ Orobanche coelestis f. typica Beck in Biblioth. Bot. 19: 114[-115] t. 2 f. 21(1) (1890) [poor quality illustration (calyx teeth seem too long)]

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? = Phelypea syspirensis K. Koch in Linnaea 22: 662 (1849) (sec. Beck, 1930: 94, "p.p."; Novopokrovskii & Tzvelev, 1958: 60; Schiman-Czeika, 1964: 9; A. Gilli, 1982: 10, "p.p."). Syn.: = Orobanche coelestis f. persica Beck in Biblioth. Bot. 19: 115 (1890) (sec. Beck, 1930: 94, "p.p.; vidi orig. in Herb. Berol."); – Orobanche coelestis sensu Novopokr. & Tzvelev in Kom., Flora SSSR 23: 60[-61] (1958). Ind. loc: [Turkey / Türkiye / Turquía, Erzurum:] "in valle Tschabantz [Tschabantz-Dereh / Çabantsdere] prope Sber [İspir] (f. 2 leg. C. Koch)" (cf. Beck, 1890: 115); [Turkey / Türkiye / Turquía:] "A8 Erzurum: Tschabanz (Çapans [Başçeşme, near İspir]) valley, 2 viii 1843, C. Koch" (sec. Gilli, 1982: 10). Obs. Status incertain. However, from our point of view, it is morphologically very different to Ph. coelestis.

? = Orobanche ehrenbergii Solms ex Beck in Engl., Pflanzenr. 96: 94 (1930) ["O. Ehrenbergii Solms daselbst"], nom. nud. Syn.: = Phelipanche coelestis (Reut.) Soják (1972) (sec. Beck, 1930: 94). Ind. loc.: "Syrien: Bei Haddet (Ehrenberg in Herb. Berlin = O. Ehrenbergii Solms daselbst)". Obs. Status incertain. However, from our point of view, it is morphologically very different to Ph. coelestis.

? = Orobanche sogdiana Novopokr. in Bot. Mat. Gerb. Bot. Inst. AN SSSR 13: 306 (1950). Syn.: ≡ Phelipanche sogdiana (Novopokr.) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972). Ind. loc.: "Gissar ridge [Qashqadaryo Province / Kashkadarya Province, Uzbekistan] (LE)" (sec. Novopokrovskii & Tzvelev, 1958: 77, 62 t. 4 f. 6). Ic.: Novopokrovskii & Tzvelev, 1958: 77, 62 t. 4 f. 6. Obs. Status incertain. However, from our point of view, it is morphologically very different to Ph. coelestis. "Parasites on Amydalus bucharica Korsh." (sec. Novopokrovskii & Tzvelev, 1958: 77)

? = Phelipanche sogdiana (Novopokr.) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972). Obs. Status incertain

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Ind. loc.: "In cacumine montis Sypili in radicibus..... (Boissier!), in Cariâ (Pinard!)." [see Mount Sipylus (Mount Spil, Manisa, Manisa-Salihli ancient capital of Lydia, Manisa Province, Turkey) and Caria (SW Turkey) in Wikipedia (accessed 17 Sept. 2007)].

Lectotypus / Lectotype: G 00147181 [herb. Prodr. (G-DC)]: A) «M Boissier 1844 / Orobanche coelestis B. & Reut.! / Caria 1843  C. Pinard»; B) «Phelipaea celestis Reut. / Orobanche celestis Boss. & Reut.». Declaramos lectótipo al ejemplar más grande de los dos del pliego.".

Designated by: Carlón, L., Gómez Casares, G., Laínz, M., Moreno Moral, G., Sánchez Pedraja, Ó. & Schneeweiss, G.M. in Documentos Jard. Bot. Atlántico (Gijón) 6: 44 (2008 [17 July 2008]).

Host: Lamiaceae[?], Phlomis sp.[?] (cf. Beck, 1890: 115, "...Haussknecht ... Phlomidis..."; Beck, 1930: 94) (cf. Annotated Checklist of Host Plants of Orobanchaceae, accessed 24 August 2009). The host plant needs to be confirmed by further studies.

Comments: "Typus: Caria, Pinard, W!" (sec. H. Schiman-Czeika, 1964: 9); "corollâ porrecta" (cf. Reuter in DC. Prod. 11: 5. 1847). O. coelestis, according to Beck 1930, is a heterogeneous mixture of different species that have little to do with each other; e.g., sec. Beck, 1930: 93-94; Schiman-Czeika, 1964: 9-10; Flora of China and Flora of Pakistan (accessed, 20 Oct. 2008) this species is conspecific with Phelipanche heldreichii (Reut.) Soják ["Gruppe der Trichocalyces (O. Schultzii, stricta, trichocalyx)" (sensu Beck, 1890: 117)]. However, from our point of view, Ph. coelestis is morphologically very different to Phelipanche heldreichii and to the "Gruppe der Trichocalyces". It is relatively morphologically similar to Phelipanche bungeana, Ph. cilicica, Ph. nowackiana (Ph. rechingeri), Ph. pulchella [incl. Orobanche longiflora sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 47, t. 695 f. 932 (1829), non Pers. (1806); – Phelypaea longiflora sensu C.A. Mey., Verz. Pfl. Casp. Meer.: 104 (1831), non Pers. (1806)].

Chromosomatic number:

Ic.: a, G 00147181 (herb. Prodr. (G-DC)]: A) «M Boissier 1844 / Orobanche coelestis B. & Reut.! / Caria 1843  C. Pinard»; lectotype); b, specimens (ib.); c, G 00074291 - Pinard 1843 (ib.; isolectotype); d, specimens (ib.); e, MA 742261 (Aragat mountain, Aragatsotn province, Armenia); f, G 00147181 (Caria; lectotype) vs. MA 742261 (Aragat).

Distribution: its distribution is not well known, because it has frequently been confused with other species (e.g., Phelipanche heldreichii [Asia, on Eryngium sp.]; Phelipanche pulchella [Asia (Imeritia, leg. Szovits)]; Ph. portoilicitana [N Africa, Bengashi, on Centaurea sp.]).

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Specimens examined and literature records.

Armenia [Caucasus / Cáucaso] (cf. Boissier: 1879: 496, sub Phelipaea celestis Reut.; Beck, 1930: 94, only f. typica)

Aragatsotn: "Iter Armeniacum (VI-VII-2005) / Orobanche / Armenia, prov. Aragatsotn, monte Aragat [Aragat mountain] / Ghazaravan, / carretera de subida al lago Kari, 40º 23' 00'' N  44º 15' 38'' E, / 2080 m, 30-jun-2005, pastos por encima del limite del / robledal, Medina & al. 2597. / Plantae Armeniae, anno 2005 ... )" (MA 742261).

Georgia [Caucasus / Cáucaso]

Imeretiya / Imereti: "f. 1—3 Szovits in itinere suo persico, quorum plantae in herb. Petrop. conservantur" (sec. Beck, 1890: 115, f. typica); "Ex herbario horti Petropolitani / Phelipaea coelestis Reut. / Imeritia [Imeretiya / Imereti]"] / Szovits [Szovits O.J.] (P 02968079 [Reçu le 16 Avril 1880]; rev. as Phelipanche pulchella by Sánchez Pedraja, 30 Nov. 2011 (cf. Reuter, 1847: 5, sub Phelypaea pulchella C.A. Mey.)); "Imeritiâ (Eichw.)" (cf. Boissier: 1879: 497, sub Ph. pulchella).

Iran / Irán [Persia]

Eşfahān / Esfahan / Isfahan: "Persia: ... in monte Taktak [Kūh-e Tak Tak / Kuh-e Taktak, N 32° 20' 23'' E 51° 41' 10'', near Mobārakeh, Eşfahān / Esfahan / Isfahan] (f. 1 leg. Haussknecht)" (sec. Beck, 1890: 115, f. typica).

Turkey / Türkiye / Turquía (Beck, 1930: 94, only f. typica).

Caria (SW Turkey): «M Boissier 1844 / Orobanche coelestis B. & Reut.! / Caria 1843  C. Pinard»; B) «Phelipaea celestis Reut. / Orobanche celestis Boss. & Reut.» (G 00147181 [herb. Prodr. G-DC], lectotypus); ibidem (G 00074291, isolectotypi); ibidem (cf. Boissier: 1879: 496, sub Phelipaea celestis Reut.).

Tokat: "prope Tokat (leg. Dr. Wiedermann in herb. Petrop.)" (sec. Beck, 1890: 115, f. typica); "In meiner Abhandlung über die Orobanchen Vorderasiens. (l.c. p. 676) ist bei den Angaben über die Verbreitung dieser Art zu ergänzen „Pontus (Tokat"), daselbst von Wiedemann (sec. Beck) gefunden. — Wie bereits der Monograph hervorhebt, steht diese Varietät von O. coelestis Boiss. et Reut. der ebenfalls bei Sultanabad [Arak / Arāk (formerly Solţānābād / Sultānābād)] is a city in and the capital of Markazi Province, Iran] vorkommenden O. Aegyptiaea ß) tricholoba (Reut.) ungemein nahe. Das vorliegende Exemplar, obwohl der Diagnose entsprechend, ist von der von G. v. Beck als tricholoba bestimmten Pflanze kaum verschieden." (cf. Bornmüller, 1907: 116, sub Orobanche coelestis Boiss. et Reut.).

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References.

Beck, G. 1890. Monographie der Gattung Orobanche. Biblioth. Bot. 19. Theodor Fischer. Kassel.

Beck, G. 1930. IV. 261. Orobanchaceae. In Engler, A. (ed.). Das Pflanzenreich. Regni Vegetabili Conspectus. Wilhelm Engelmann. Leipzig [pp. 1-348].

Boissier, P.É. 1879 [Apr.-May 1879]. Flora orientalis [...] Volumen quartum. Fasciculus secundus. Genevae et Basileae. [Ord. LXXXIX. Orobanchaceae: 492-518].

Bornmüller, J. 1907. Plantae Straussianae sive enumeratio plantarum a Th. Strauss annis 1889-1899 in Persia occidentali collectarum. Pars III. Beih. Bot. Centralbl. Abt. 2, 22: 102-142 (accessed, 23 March 2011).

Carlón, L., Gómez Casares, G., Laínz, M., Moreno Moral, G., Sánchez Pedraja, Ó. & Schneeweiss, G.M. 2005 [June 2005]. Más, a propósito de algunas Orobanche L. y Phelipanche Pomel (Orobanchaceae) del oeste del Paleártico [More on some Orobanche L. and Phelipanche Pomel (Orobanchaceae) from the western Palearctic]. Documentos Jard. Bot. Atlántico (Gijón) 3: 1-71 (in Spanish, plus an Appendix in English). [Texto completo (pdf) / Full Text (pdf)] [Suplemento (pdf) / Supplement (pdf)] [Índice (htm) / Index (html) 3].

Carlón, L., Gómez Casares, G., Laínz, M., Moreno Moral, G., Sánchez Pedraja, Ó. & Schneeweiss, G.M. 2008 [17 July 2008]. Más, a propósito de algunas Phelipanche Pomel, Boulardia F. W. Schultz y Orobanche L. (Orobanchaceae) del oeste del Paleártico [More on some Phelipanche Pomel, Boulardia F.W. Schultz and Orobanche L. (Orobanchaceae) from the western Palearctic.]. Documentos Jard. Bot. Atlántico (Gijón) 6: 1-128 (in Spanish, plus an enlarged abstract in English). [Texto completo (pdf) / Full Text (pdf) in GBC and Texto completo (pdf) / Full Text (pdf) in Biblioteca Digital del Real Jardín Botánico].

Edmondson, J. R. & Lack, H.W. 2006. Karl Georg Theodor Kotschy’s itinerary in southern Iran, 1841-42. Willdenowia 36 (Special Issue): 579-588 (pdf).

Gilli, A. 1982. Orobanchaceae (90). In Davis, P.H. (ed.). Flora of Turkey 7: 1-22. Edinburgh.

Flora of China - Orobanchaceae [eFloras.org - http://www.efloras.org/] (accessed 11 Sept. 2007).

Flora of Pakistan Home Page - Flora of Pakistan Online Version - Orobanchaceae [eFloras.org - http://www.efloras.org/] (accessed 11 Sept. 2007).

Greuter, W., Burdet, H.M. & Long, G. 1989. Med-Checklist.. 4: 265-266 [Orobanchaceae]. Genève, Berlin.

Koch, K. 1846. Wanderungen im Oriente, während der Jahre 1843 und 1844. Reise im pontischen Gebirge und türkischen Armenien. (pdf).

Koch, K. 1849. Beiträge zu einer Flora des Orientes. Linnaea 22: 177-338, [Fortsetzung] 597-752. [Orobanchaceae: 658-670].

Novopokrovskii, I.V. & Tzvelev, N.N. 1958. Orobanchaceae Vent. In Komarov, V. L., (Ch. Ed.). 1934-1964. Flora SSSR 23 : [I]-XXIII, [1]-775. Edit. Akad. Nauk SSSR. Moskva-Leningrad. [Panigrahi, G. (Bot. Ed.). 2000. Bishen Singh Mahendra Pal Singh].

Reuter, G. 1847. Orobanchaceae. In De Candolle, Alphonse (ed.). Prodromus Systematis naturalis regni vegetabilis... 11: 1-45. [Published on 25 Nov. 1847].

Schiman-Czeika, H. 1964. Orobanchaceae. In Rechinger, K.H. (ed.). FLora Iranica. Graz-Austria. N.º 5 [15 July 1964].

Soják, J. 1972. Nomenklatorické Poznámky (Phanerogamae). Cas. Nár. Mus., Odd. Prír. 140: 127-134.

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