Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

ISSN: 2386-9666 - Last Updated: 26 November 2025

Accepted Name:

Phelipanche arenaria (Borkh.) Pomel in Bull. Soc. Sci. Phys. Algérie 11: 103 (1874) [Nouv. Mat. Fl. Atl. 11: 103 (1874 [post 10 Mar 1874])]

Basionymum / Basionym:

≡ Orobanche arenaria Borkh. in Neues Mag. Bot. 1: 6 (1794)

Synonyms:

Homotypic synonyms:

≡ Kopsia arenaria (Borkh.) Dumort., Comment. Bot.: 17 (1822), nom. rejic. (ICNB - Tokyo Code, Index Kopsia - 6626 Angiospermae, Dicotyledones, Metachlamydeae 6165-9629, accessed Feb 2006; Vienna Code, 2006 Appendix III Nomina generica conservanda et rejicienda - E3. Dicotyledones, accessed Jun 2009)

≡ Phelypaea arenaria (Borkh.) Walp., Repert. Bot. Syst. 3: 459 (1844) ["Phelipaea"]

≡ Lathraea arenaria (Borkh.) Cuatrec. in Treb. Mus. Ci. Nat. Barcelona 12, Sèr. Bot.: 429 (1929) ["Lathraea arenaria; Phelipaea arenaria Walp."]

≡ Orobanche arenaria f. typica Beck in Biblioth. Bot. 19: 129 (1890) ["Formas ... 1. typica = Orobanche arenaria Borkhausen."], nom. inval. [not validly published] (Turland & al. 2018: Art. 24.3)

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Heterotypic synonyms:

= Orobanche laevis L., Sp. Pl. 2: 632 (1753), nom. rej. Syn.: ≡ Phelipanche laevis (L.) Holub, Preslia 62: 196 (1990); = Phelipanche purpurea (Jacq.) Soják (1972) [p.p.]; = Phelipanche arenaria (Borkh.) Pomel (1874) [p.p.]. Ind. loc.: France: "Habit Monspelii [Montpellier, Hérault]". Obs. "ambiguous name" [nom. rej. prop. (Turland & Rumsey in Taxon 46: 789-791. 1997); nom. rej. (Brummitt in Taxon 49: 263-264. 2000)]

≠ Orobanche longiflora Trevir. in Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 7: 150 t. 2 f. 10, 11 (1816) [nom. illeg.], non Orobanche longiflora Pers., Syn. Pl. 2(1): 181 (1806). Syn.: ≡ Phelypaea longiflora (Trevir.) G. Don, Gen. Hist. 4: 632 (1838) [nom. illeg.]; = Orobanche aegyptiaca var. typica f. rectiflora Beck in Biblioth. Bot. 19: 102 (1890) (Beck 1890: 102); = Orobanche longiflora sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 47, t. 695 f. 932 (1829), non Pers. (1806); = Kopsia longiflora sensu Dumort., Comment. Bot.: 18 (1822), non Pers. (1806); = Phelypaea longiflora sensu C.A. Mey., Verz. Pfl. Casp. Meer.: 104 (1831), non Pers. (1806). Ind. loc.: Russia: "Sareptae leg. ill. Erdmann" [Old Sarepta / Старая Сарепта (now part of Volgograd), Southeast of the European part of Russia / Россия] (Reichenbach 1829: 47, "Treviran. Act. Berol. 1813. II. t. 2. f. 10."); "H. Native of Siberia, at the Volga" (Don 1838: 632). Herbarium Specimens: 1. "Phelipaea longiflora C.A.M. / ad oppid. Guder [?]". – 2. "Имперaторckаго Моcковскаго Университета [Imperial Moscow University] / Gremiatschensky, S.J. Die Pflanzen der Bukejevvschen / Kirgisen-Horde. 1855. / (Ryn-Peski Sanddünen [Ryn Desert / Рын-пески / Нарын-Құм / Naryn-Qumt], Gross. Bogdo, Bisch-tau, Jakschibisch-tau Arsagar, Baskuntschatskoje-See [Astrakhan steppe / Kalmyk Steppe] et сеt.)". – 3. "Orobanche arenaria Borkh. / 29.VIII.1949 / Det. I. Novopokrovsky" (https://plant.depo.msu.ru/open/public/item/MW0514043 - https://plant.depo.msu.ru/open/public/item/MW0514043/img/0.jpg?original=2904dadb-71b9-441d-a832-aa8098839291 - MW 0514043.jpg - https://www.gbif.org/occurrence/1697663136; rev. Ó. Sánchez Pedraja May 2024, sub P. arenaria). Obs. ? = Phelipanche arenaria (Borkh.) Pomel (1874) [Orobanche longiflora sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 47, t. 695 f. 932. 1829 (Reichenbach f. 1862: 86, "Forsan etiam Orobanche longiflora P. Rchb. Crit. VII. DCXCV")]; ? = Phelipanche pulchella (C.A. Mey.) Soják (1972) (Greuter & al. 1989: 256), this species is treated by Greuter & al. (1989) as conespecific with Orobanche aegyptiaca, but according to Novopokrovskii & Tzvelev (1958[2000]: 54 t. 4 f. 1) it is a different species. See Phelipanche aegyptiaca (Pers.) Pomel

= Orobanche comosa Wallr., Sched. Crit. 1: 314 (1822) (Walpers 1844: 460). Syn.: ≡ Phelypaea comosa (Wallr.) G. Don, Gen. Hist. 4: 632 (1838 [8 Mar-8 Apr 1838]); ? = Orobanche vagabonde Vaucher (1827), nom. inval. (Mutel 1835: 352); = Phelipanche arenaria (Borkh.) Pomel (1874) (Reichenbach 1829: 46; Schultz 1830: 493, "?"; Mutel 1835: 352; Walpers 1844: 460; Beck 1890: 129 "vidi orig. in herb. Berol. [BW 11593030]"; Beck 1930: 102). Ind. loc.: Germany: "ad Genistae tinctoria [mistake] radices in colle herbido der Steinklippe. Jun. [near Wendelstein, SE of Rossleben / Roßleben (Until 1815 Roßleben was located in Wendelstein, Saxony), Thüringen, Germany / Deutschland]" (Skočdopolová & Chrtek 2008: 26). Obs. "Herba viva pallide ... Spica comosa ... condensata ... Corolla ... crenulatum ... Calyx ...connatus, quinquefidus ... Antherae ... ciliatae" (Wallroth, l.c.). "The specimen PR 11757 is selected here as a lectotype. According to the January 2000 revision by German botanist J. Pusch this specimen corresponds to Orobanche arenaria Borkh." (Skočdopolová & Chrtek 2008: 26). Herbarium Specimen: 1. "O. arenaria Borkh. / Beck [m. Beck]. – 2. "O. comosa mea". – 3. "187 W" (BW 11593030; rev. Ó. Sánchez Pedraja Sep 2016)

= Orobanche de l'Artemise commune Vaucher, Monogr. Orobanch.: 65 [t. 14] (1827) ["XXVII ... Planche 14"], excl. syn., nom. inval. [not validly published]. Syn.: ≡ Orobanche artemisia-vulgaris Duby, Bot. Gall.: 350 (1828), pro syn.; ≡ Orobanche artemisiae-vulgaris F.W. Schultz in Ann. Gew. Regensb. 5: 500 (1830) ["27. O. Artemisiae vulgaris Tab. XIV"] pro syn.; = Orobanche caerulea Vill. (1787) (Duby 1828: 350; Schultz 1830: 500-501; Mutel 1835: 352); = Orobanche purpurea Jacq. (Vaucher 1827: 65; Reichenbach 1829: 46; Schultz 1830: 500); = Orobanche arenaria Borkh. (Reichenbach 1829: 46; Beck 1890: 129, "exclus. Synon. et tab. 14"). Ind. loc.: "… dans presque toute l’Europe. … 1825 et 1826, sur les collines qui bordent la route de Saint-Pierre à Sion [Rhône-Alpes, France], à demi-lieue de cette dernière ville. Elle est exclusivement parasite de l’Artémise vulgaire, comme l’indique aussi Wallroth…". Obs. "anthères … cotonneuses" (Vaucher 1827: 65); = Orobanche caerulea DC. ... Wiild. ... Wallr. et = Orobanche purpurea Jacq. (Vaucher 1827: 65)

= Orobanche artemisia-vulgaris Duby, Bot. Gall.: 350 (1828), pro syn. ("cited as a synonym", Turland & al. 2018:  Art. 52.1 et Art. 52.2). Syn.: ≡ Orobanche de l'Artemise commune Vaucher, Monogr. Orobanch.: 65 [t. 14] (1827), nom. inval.

= Orobanche artemisiae-vulgaris F.W. Schultz in Ann. Gew. Regensb. 5: 500 (1830) ["27. O. Artemisiae vulgaris Tab. XIV"], pro syn. Syn.: ≡ Orobanche de l'Artemise commune Vaucher, Monogr. Orobanch.: 65 [t. 14] (1827), nom. inval.)

= Orobanche borkhausenii Andrz. ex Besser in Flora 15(2): 19 (1832), nom. nud. (Beck 1890: 129, "Orobanche Borkhausenii Andrz. ex Besser, Flora (1832) II, Beibl. p. 19, fide Ledebour. Vidi orig. c. d. Besseri in herb. Petropol. et in herb. Vindob. [collect. Reich. fil.]"; Beck 1930: 102, "O. Borkhausenii Andrz. ex Besser in Flora II. (1832) Beibl. 19, fide Ledebour (vidi spec. orig. d. Besseri conservata in herb. Vindob. et Petropol.)"). Syn.: ≡ Orobanche borkhausenii Besser ex Reut. in DC., Prod. 11: 6 (1847); ≡ Kopsia borkhausenii (Besser ex Reut.) Caruel in Parl., Fl. Ital. 6: 355 (1885). Ind. loc.: Ukraine: "in Podolien ... die Andrzejowski Borkhausenii nannte" (Besser 1832: 19). "Podolia [Podolia / Поділля / Podillia / Подолье / Podolye, Vinnytsia Oblast / Вінницька область / Vinnyts’ka oblast’] Besser!" [?] (Reichenbach f. 1862: 87, sub P. arenaria Walp. [P. robusta Walp.]). Herbarium Specimens: «Authentic specimen ["Original material"]: A) ‘Herb. Ledebour // Phelipaea arenaria Walpers // Orobanche borkhausenii / Nobis / O. arenaria / Prim[itiae]. Fl[orae]. Gal[iciae (Galicia / Galizia / Galicja / Галичина) )]. / In nemorosis. / Herb. W. Besser.’ B) ‘Herb. Ledeb. 757.b. / Philipaea arenaria Walp.’ C) ‘Orobanche arenaria Borkh. / Подолия (Сравн. Ledeb. Fl. Ross. III, 312) / 25.IX.1947 Determ. I. Novopokrovsky’ D) ‘Lectotypus Orobanche / borkhausenii Andrz. ex Besser / Holger Uhlich, Welterod, Germany 2020’ (LE 01060137 [http://re.herbariumle.ru/01060137 - LE 01060137.jpg], Supporting information)» (Uhlich & Rätzel 2024: 6, sub P. arenaria); Type?: "Orobanche Borkhausenii / Nobis / O arenaria Prim Agah [?] / In nemorosis / Herb. W. Besser / III 4" (K 000759344 - Herb. W. Besser - http://specimens.kew.org/herbarium/K000759344 - http://www.kew.org/herbcatimg/400395.jpg; rev. Ó. Sánchez Pedraja Mar 2012 [the sheet contains four specimens (each of them next to its label), three of them correspond to P. purpurea from Jersey (specimens of Lagasga [K 000759342 - hb. Gay] and Stevens [K 000759343 - hb. Stevens]) and the fourth (the Besser's specimen) seems to correspond to P. arenaria from Podolien?]). Obs. Sinónimos de Orobanche arenaria Borkh. (Flora iberica, accessed Oct 2007)

= Orobanche robusta A. Dietr., Fl. Boruss. 3: 151 [t. 151] (1835). Syn.: ≡ Phelypaea robusta (A. Dietr.) Walp., Repert. Bot. Syst. 3: 460 (1844); ≡ Orobanche arenaria f. robusta (A. Dietr.) Beck in Biblioth. Bot. 19: 130 (1890). Ind. loc.: Germany: "Auf sonnigen Auhöhen im Sandboden bei Baumgartenbrück hinter Potsdam [Brandenburg, Germany / Deutschland] im Jahr 1834 in Gemeinschaft mit dem Herrn Cabinets-Secretair Sasse entdeckt, wo sie einzeln und in Büschen wahrscheinlich auf Artemisia campestris wächst*)" (Dietrich 1835: 151); "Crescit in radicibus Artemisiae campestris in collibus arenosis apricis (bei Baumgartenbrück inter Potsdam)" (Walpers 1844: 460); "Auf sonnigen Anhöhen im Sandboden bei Baumgartenbrück inter Potsdam im Jarhr 1834 in Gemeinschaft mit dem Herrn Cabinets … und in Büschen wahrscheinlich auf Artemisia campestris wächst. Blüht im Juli" (Reuter 1847: 9, sub Phelypaea robusta, "Borussiae prope Potsdam")

= Phelypaea comosa (Wallr.) G. Don, Gen. Hist. 4: 632 (1838 [8 Mar-8 Apr 1838]) ["Phelipaea"]. Syn.: ≡ Orobanche comosa Wallr., Sched. Crit. 1: 314 (1822) (Walpers 1844: 460; Beck 1930: 102; IPNI); = Orobanche vagabonde Vaucher (1827) (Don 1838: 632)

? = Phelypaea longiflora (Trevir.) G. Don, Gen. Hist. 4: 632 (1838) [nom. illeg.]. Syn.: ≡ Orobanche longiflora Trevir. in Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 7: 150 t. 2 f. 10, 11 (1816) [nom. illeg.], non Orobanche longiflora Pers., Syn. Pl. 2(1): 181 (1806). Obs. see Phelipanche cilicica (Reut.) Soják (1972)

= Phelypaea robusta (A. Dietr.) Walp., Repert. Bot. Syst. 3: 460 (1844) ["Phelipaea"] (Walpers 1844: 460). Syn.: ≡ Orobanche robusta A. Dietr., Fl. Boruss. 3: 151 [t. 151] (1835)

= Orobanche borkhausenii Besser ex Reut. in DC., Prod. 11: 6 (1847) ["Orobanche Borkhauseni Bess.!"], pro syn. ("cited as a synonym", Turland & al. 2018:  Art. 52.1 et Art. 52.2). Syn.: ≡ Orobanche borkhausenii Andrz. ex Besser in Flora 15(2): 19 (1832), nom. nud.; ≡ Kopsia borkhausenii (Besser ex Reut.) Caruel in Parl., Fl. Ital. 6: 355 (1885) ["Borkhauseni"], nom. nud. (Caruel 1847: 356, "Bess. sec. Benth. in Cand. prodr. 11. p. 6.")

= Orobanche abortiva Leyss. ex Reut. in DC., Prod. 11: 37 (1847), pro syn. Syn.: = Phelypaea arenaria (Borkh.) Walp., Repert. Bot. Syst. 3: 459 (1844) (Walpers 1844: 460, "Orchis abortiva Leyss."; Reuter 1847: 37)

≠ Phelypaea obtusiloba Hausskn., Iter Syr. Arm. Exsicc. (1865), p.p. (Boissier 1879: 496). Syn.: = Phelipanche iberica (Beck) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972)

= Orobanche ionantha A. Kern. in Oesterr. Bot. Z. 24: 46[236] (1874). Syn.: ≡ Orobanche arenaria f. ionantha (A. Kern.) Beck in Biblioth. Bot. 19: 130 (1890) (Beck 1930: 105, "vidi orig. in herb. Kerner"); ≡ Kopsia arenaria f. ionantha (A. Kern.) Bég. in Fiori & Paol., Fl. Italia 2: 473 (1902); = Orobanche arenaria var. colorata Grüner in Bull. Soc. Nat. Mosc. 1: 91 (1869) (Beck 1890: 130; Beck 1930: 105). Ind. loc.: "Unter den in Ungarn [Hungary] gesammelten Orobanchen fand sich auch ein Exemplar einer noch nicht beschriebenen Art, welche ich seither auch an mehreren Punkten in den Thälern der Alpen auf Artemisia campestris auffand, und die ich insbesonders in der Nähe von Innsbruck [Tyrol, Austria] auf den Angern der sogenannten "Höttinger Aue [Höttinger Au]"..." (Kerner 1874: 46). Herbarium Specimens: 1. "Orobanche ionantha K. / Tirol. Trautson bei Matrei [Matrei in Osttirol (Lienz District)] / Auf Artemisia campestris / Kerner". – 2. Syntypus WU 0076573 / Orobanche ionantha A. Kern. / ... / typifcavit M. Lachmayer (WU) 2014-05-21 / ..." (https://wu.jacq.org/WU0076573 - https://www.jacq.org/detail.php?ID=482238 - WU 0076573.jpg - https://object.jacq.org/europeana/WU/482238.jpg - https://www.gbif.org/occurrence/1830990375; rev. Ó. Sánchez Pedraja May 2024, syntypes of O. ionantha); "Dr. C. Baenitz, Herbarium Europaeum Nº ... / Orobanche ionantha A. Kern. / O. cerulea Vill proparte / Tirol: Ad radices Artem. camp. circa / Innsbruck, raro 1882.VI / leg. J. Murr." (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04419637 - P 04419637.jpg - http://www.gbif.org/occurrence/465125215 - https://explore.recolnat.org/occurrence/779BED04FAD54B019FDCBF765B6420AA; rev. Ó. Sánchez Pedraja May 2018); idem (LY 0737615.jpg - hb. Rouy in Roland Bonaparte - https://www.gbif.org/occurrence/3427999059 - https://explore.recolnat.org/occurrence/C44CE22AE4804B30A725E257ADD29951; rev. Ó. Sánchez Pedraja May 2024); 1. "Orobanche arenaria / Auf Artemisia campestris. / Wald Adamow in Oberneutra [Adamovský les wood near the village of Gbely, Horná Nitra] / Juli 1853 / J.F Kržišch". – 2. "Orobanche ionantha K. / Kerner". – 3. "... O. ionantha ..." . – 4. "Syntypus / Orobanche ionantha A. Kern. / Oesterr. Bot. Z. 24: 46-47 (1874) / typificavit M. Lachmayer (WU) 2014-05-21 /..." (WU 0076578 - https://wu.jacq.org/WU0076578 - https://www.jacq.org/detail.php?ID=482230 - https://www.gbif.org/occurrence/1830990253; rev. Ó. Sánchez Pedraja Jul 2022)

≠ Phelypaea atlantica Pomel in Bull. Soc. Sci. Phys. Algérie 11: 103 (1874) [Nouv. Mat. Fl. Atl. 11: 103 (1874 [post 10 Mar 1874])] ["Phelipaea atlantica Pom. h." (in sched.)], nom. illeg. Syn.: ≡ Phelipanche atlantica Pomel in Bull. Soc. Sci. Phys. Algérie 11: 103[-104] (1874); = Phelipanche purpurea subsp. ballii (Maire) Carlón, G. Gómez, M. Laínz, Moreno Mor., Ó. Sánchez & Schneew. in Documentos Jard. Bot. Atlántico (Gijón) 6: 104 (2008 [17 Jul 2008])

≠ Phelipanche atlantica Pomel in Bull. Soc. Sci. Phys. Algérie 11: 103[-104] (1874) [Nouv. Mat. Fl. Atl. 11: 103[-104] (1874 [post 10 Mar 1874])] (Battandier 1890: 656, sub Ph. arenaria Walpers). Syn.: ≡ Parabanche caerulea var. atlantica Pomel (1860 [24 May 1860]), in sched., nom. nud.; ≡ Phelypaea atlantica Pomel in Bull. Soc. Sci. Phys. Algérie 11: 103 (1874), nom. illeg.; = Phelipanche purpurea subsp. ballii (Maire) Carlón, G. Gómez, M. Laínz, Moreno Mor., Ó. Sánchez & Schneew. in Documentos Jard. Bot. Atlántico (Gijón) 6: 104 (2008 [17 July 2008])? = Orobanche arenaria f. stenosepala Beck in Biblioth. Bot. 19: 130 (1890) (Beck 1930: 105). Ind. loc.: Algeria: "Sur Artemisia campestris: Mou-el-Gtouta." [Hauts plateaux Oranais, Oran, Algérie / Algeria / Argelia]. Herbarium Specimens: 1. "Parabanche caerulea / var. atlantica / itima 25 mai 60 [m. Pomel]". – 2. "... Orobanche laevis L. / = Phelipanche atlantica Pomel! (Type!) / O. Itima, Mou-el-Gtouta [au S. de Tiaret, Hauts plateaux Oranais, Oran department] / 25-5-1860  leg. A. Pomel [m. Maire]" (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu008101 - MPU 008101.jpg - herb. Pomel - http://www.gbif.org/occurrence/1424544992; rev. Ó. Sánchez Pedraja Nov 2009, holotype of Phelipanche atlantica [= Phelipanche purpurea subsp. ballii])

≠ Phelypaea obtusiloba Reut. ex Boiss., Fl. Orient. 4(2): 496 (1879 [Apr-May 1879]) ["Phelipaea obtusiloba Reut. in Haussk. exs."], pro syn. Syn.: ≡ Phelypaea obtusiloba Hausskn., Iter Syr. Arm. Exsicc. (1865) (Beck 1890: 126); – Orobanche purpurea var. iberica Beck in Biblioth. Bot. 19: 126 (1890), p.p. (Beck 1890: 126). Ind. loc.: Turkey: "in monte Berytdagh Cataoniae [Berit Dağı, Kahramanmaraş Province] 8000' (Haussk!)" (Boissier 1879: 496). Obs. view IPNI entry

? = Orobanche pareysi Beck in Oesterr. Bot. Z. 31: 310 (1881). Syn.: ≡ Phelypaea pareysii (Beck) Boiss., Fl. Orient., Suppl.: 357 (1888 [Oct 1888]); ≡ Orobanche purpurea var. pareysi (Beck) Beck in Biblioth. Bot. 19: 127 (1890). Ind. loc.: Krim: "Habitat: In Tauria ubi legit cl. dom. Pareys, cujus specimen in herbario caesareo Vindobonensi conservatur" (Beck 1881: 310); "Hab. Tauria (Pareysz in Herb. Caes. Vindob.) (Non vidi)" (Boissier 1888: 357); "Tauria! (herb. Vindob.)" (Beck 1890: 127); "Russische Länder: In der Krim (Pareys in Herb. Wien!)" (Beck 1930: 112). Obs. "apparently is described from random, not fully typical specimens, possibly also belonging to O. arenaria Borkh. Beck reported it for Crimea and Talysh [Iran]" (Novopokrovskij & Tzvelev 1958: 45[42], sub O. purpurea)

= Kopsia borkhausenii (Besser ex Reut.) Caruel in Parl., Fl. Ital. 6: 355 (1885) ["Borkhauseni"], nom. inval. (Caruel 1847: 356, "Bess. sec. Benth. in Cand. prodr. 11. p. 6."). Syn.: ≡ Orobanche borkhausenii Besser ex Reut. in DC., Prod. 11: 6 (1847)

≠ Orobanche iberica (Beck) Tzvelev in Karjagin, Fl. Azerbaĭdzhana 7: 566 (1957) (Uhlich & Rätzel 2024: 19, sub P. purpurea var. iberica). Syn.: ≡ Phelipanche iberica (Beck) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972); ≡ Orobanche purpurea var. iberica Beck, Biblioth. Bot. 19: 126 (1890)

≠ Orobanche iberica (Beck) Tzvelev in V.L. Komarov, Fl. URSS 23: 49 (1958), nom. superfl. (Uhlich & Rätzel 2024: 19, sub P. purpurea var. iberica). Syn.: ≡ Phelipanche iberica (Beck) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972); ≡ Orobanche purpurea var. iberica Beck, Biblioth. Bot. 19: 126 (1890)

≠ Phelipanche iberica (Beck) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972) ["cf. iberica"] (Uhlich & Rätzel 2024: 19, sub P. purpurea var. iberica). Syn.: ≡ Orobanche purpurea var. iberica Beck, Biblioth. Bot. 19: 126 (1890)

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= Phelypaea arenaria var. subramosa Noulet, Fl. Bass. Sous-Pyren.: 489 (1837) ["6. Un peu rameuse. Subramosa. Nob."]. Syn.: = Orobanche arenaria f. robusta (A. Dietr.) Beck in Biblioth. Bot. 19: 130 (1890) (Beck 1930: 104). Ind. loc.: France: "Flore du bassin sous-pyrénéen ... aux environs de Toulouse [Haute-Garonne, Midi-Pyrénées]"

? = Phelypaea arenaria var. rubentiarenaria Reut. in DC., Prod. 11: 6[-7] (1847). Ind. loc.: France: "Ad radices Medicaginis prope Lugdunum [Lyon] (Al. Jordan!) ... (v. in h. Jordan.)"

= Phelypaea arenaria var. elongata Koch in Linnaea 22: 661 (1849) (Beck 1930: 102, "vidi origin.!"). Ind. loc.:  Armenia?: "In der Ebene des Araxes [Aras river, Caucasus], c. 2700' hoch" (Koch 1849: 661). Obs. "Unterscheidet sich durch die Grösse von der Hauptort, aber auch dadurch, dass der Griffel die Antheren überragt" (Koch 1849: 661)

= Phelypaea arenaria var. campylantha Rchb. f., Icon. Fl. Germ. Helv. 20: 87 [t. 146] (1862 [22 Dec 1862]) (Beck 1930: 102). Ind. loc.: "Habui pro specie, sed characterem energicum frustra quacsivi."

= Orobanche arenaria var. colorata Grüner in Bull. Soc. Nat. Mosc. 1: 91 (1869) ["β"]. Syn.: = Orobanche ionantha A. Kern. in Oesterr. Bot. Z. 24: 46[236] (1874) (Beck 1890: 130)

≠ Orobanche purpurea var. iberica Beck, Biblioth. Bot. 19: 126 (1890) [“Variat: … ζ . (Orobanche) iberica”]. Syn.: ≡ Orobanche iberica (Beck) Tzvelev in Karjagin, Fl. Azerbaĭdzhana 7: 566 (1957); ≡ Orobanche iberica (Beck) Tzvelev in V.L. Komarov, Fl. URSS 23: 49 (1958), nom. superfl.; ≡ Phelipanche iberica (Beck) Soják in Čas. Nár. Muz. Přir. 140: 130 (1972); = Phelypaea obtusiloba Hausskn., Iter Syr. Arm. Exsicc. (1865), p.p. (Beck 1890: 126); = Phelypaea obtusiloba Reut. ex Boiss., Fl. Orient. 4(2): 496 (1879 [Apr-May 1879]), pro syn. Ind. loc.: Turkey: "Vidi ex Iberia, ex monte Berytdagh Cataloniae [Beryt Dagh, Cataonia (Cappadocia)] (leg. Haussknecht) etiam ex Germania formas similis." (Beck 1890: 126). Herbarium Specimens: "Lectotype (designated here): A) ‘Professor C. Haussknecht. Iter Orientale 1865. Phelipaea obtusiloba Hskn. et Reut. Ph. arenaria Boiss. Fl. Or. In silvis Populi tremuli in Mte. Berytdagh Cataoniae 7000’, dieb. 9./8. [1865]’. B) ‘Orobanche iberica m[ihi]. {O. purpureae forma} det. Dr G. Beck’ (JE00029051; Supporting information). The herbarium specimen from Vienna (W0171551, collected 8 Aug. 1865) is a syntype" (Uhlich & Rätzel 2024: 19, sub P. purpurea var. iberica). Obs. "Parasites on ... Artemisia Meyeriana Bess. and others ... Type: Described from Turkey (W) ... much closely to O. arenaria Borkh."; https://www.ipni.org/n/662840-1; Carlón & al. 2005: 60 f. 3 ["Phylogenetic relationships"], 66, sub "cf. iberica"; Carlón & al. 2008: 12 f. 1 [phylogenetic relationships] sub "cf. iberica"

= Phelypaea arenaria var. euxina Velen., Fl. Bulg. Suppl. 1: 222 (1898). Syn.: ≡ Orobanche arenaria var. euxina (Velen.) D. Delip., Fl. Republ. Bŭlgariya 10: 302 (1995) (https://www.ipni.org/n/992028-1). Ind. loc.: Bulgaria: "Pontum [Pontus Euxinus / Black Sea (Varna)] ‭(Str) ..." (Velenovský 1898: 222, sub Phelipaea arenaria); "Varna (Střibrny)" (Beck 1930: 103, sub O. arenaria). Herbarium Specimens: 1. "Flora bulgarica / Phelypaea arenaria Walp. ! Beck [m. Beck] / β) euxina m. Varna (Stribrny) / 1897.V Legit [det.] dr. J. Velenovský". – 2. "Orobanche arenaria Borkh. / 1970 Revidit: K. Kult". – 3. "Typus" (PRC 451846 - https://prc.jacq.org/PRC451846 - https://www.jacq.org/detail.php?ID=559522 - https://www.gbif.org/occurrence/3413676613 - http://www.jacq.org/image.php?filename=559522&method=europeana; rev. Ó. Sánchez Pedraja Jul 2022)

= Phelypaea arenaria var. ramosa Merino [before 1906, in sched., nom. nud.] ["Phelipaea"]. Syn.: ≡ Phelypaea arenaria f. ramosa [Merino ex] M. Laínz in Brotéria Ci. Nat. 24(51): 125[18] (1955). Ind. loc.: Spain: "En los primeros arenales del Miño [Pontevedra, Galicia] (MA 114834!)". Obs. "Vive a orillas del Miño cerca de su desembocadura á expensas de la Artemisia crithmifolia L. [= A. campestris subsp. maritima Arcang., Compositae] Fl. por Junio (V. vivam). ... existen mezclados pies de tallos ramosos con otros que le tienen sencillo" (Merino, Fl. Galicia 2: 53-54. 1906)

= Phelypaea caesia [var.] ampliflora Pau ["Phelipaea caesia ?ampliflora? ... [description]", in sched., nom. inval. Syn.: ≡ Phelypaea arenaria var. ampliflora [Pau ex] Cuatrecasas [in sched., nom. inval.]. Ind. loc.: Spain: "In monte "La Rapita" Sierra de Espadán [Castellón / Castelló, Valencian Community], 29 Junii 1902 [m. C. Pau]" (MA 435485).

= Phelypaea arenaria var. ampliflora [Pau ex] Cuatrecasas [in sched., nom. nud.] (MA 114832)

= Orobanche arenaria var. euxina (Velen.) D. Delip., Fl. Republ. Bŭlgariya 10: 302 (1995) (https://www.ipni.org/n/992028-1). Syn.: ≡ Phelypaea arenaria var. euxina Velen., Fl. Bulg. Suppl. 1: 222 (1898)

= Phelipanche arenaria var. personata (Beck) Uhlich & Rätzel in Nordic J. Bot. e04155: 5 (2024). Syn.: ≡ Orobanche arenaria f. personata Beck in Repert. Spec. Nov. Regni Veg. 18: 35[467] (1922)

= Phelipanche arenaria var. sareptana (Beck) Uhlich & Rätzel in Nordic J. Bot. e04155: 6 (2024). Syn.: ≡ Orobanche arenaria f. sareptana Beck in Biblioth. Bot. 19: 130 (1890)

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= Orobanche arenaria f. gigantiflora Borbás in Homokpuszt. Növ. 106 (1886) (Beck 1930: 105). Ind. loc.: "In Ungarn [Hungary!], Spanien [Spain] gefunden" (Beck 1930: 105)

= Orobanche arenaria f. ionantha (A. Kern.) Beck in Biblioth. Bot. 19: 130 (1890) ["Formas ... 2. (Orobanche) ionantha A. Kerner"] (Beck 1930: 105). Syn.: ≡ Orobanche ionantha A. Kern. in Oesterr. Bot. Z. 24: 46[236] (1874)

= Orobanche arenaria f. lamprantha Beck in Biblioth. Bot. 19: 130 (1890)

= Orobanche arenaria f. leptocaulon Beck in Biblioth. Bot. 19: 130 (1890)

= Orobanche arenaria f. prominens Beck in Biblioth. Bot. 19: 130 (1890)

= Orobanche arenaria f. robusta (A. Dietr.) Beck in Biblioth. Bot. 19: 130 (1890) ["Formas ... 4. (Orobanche) robusta Dietrich"]. Syn.: ≡ Orobanche robusta A. Dietr., Fl. Boruss. 3: 151 [t. 151] (1835) (Beck 1930: 104; Novopokrovskii & Tzvelev 1958: 49 [58])

= Orobanche arenaria f. sareptana Beck in Biblioth. Bot. 19: 130 (1890) ["Formas ... 6. (Orobanche) Sareptana"]. Syn.: ≡ Phelipanche arenaria var. sareptana (Beck) Uhlich & Rätzel in Nordic J. Bot. e04155: 6 (2024). Ind. loc.: "Vidi specimina tria, primum Sareptanum [Old Sarepta / Старая Сарепта, Volgograd, Volgograd Oblast, Russia] in herb. Petrop., secundum a cl. d. Winter prope Aranjuez in Hispania [Madrid, Spain / España], tertium prope St. Ander [Santander, Cantabria, Spain / España] a cl. Levier collectum.". Herbarium Specimens: «Lectotype (designated here): A) ‘Sarepta [Old Sarepta / Старая Сарепта] / Herb. Fischer.’ B) ‘O. arenaria Borkh. / f. O. sareptana / Beck’ [manu Beck] C) ‘O. arenaria Borkh. / (из Сарепты) / 7 V 1955 Teste: N. Tzvelev’ [manu Tzvelev] D) ‘Syntypus Orobanche arenaria / f. sareptana Beck 1890 / Holger Uhlich, Welterod, Germany 2020’ (LE 01060142 [http://re.herbariumle.ru/01060142 - LE 01060142.jpg], Supporting information)» (Uhlich & Rätzel 2024: 6, sub P. arenaria var. sareptana). Types: LE ?. "Phelipaea lanuginosa C.A. Mey. / Rossia australis: Sarepta [Old Sarepta / Старая Сарепта] Locis arenosis / montanis  M. Junii / 1883.  leg. A. Becker" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04409198 - P 04409198.jpg - http://www.gbif.org/occurrence/464896678 - https://explore.recolnat.org/occurrence/A9E50E66EE0143C991B7450152F867D1; rev. Ó. Sánchez Pedraja Nov 2011 [the sheet contains two specimens which seem to correspond to P. arenaria]); "Phelipaea arenaria / [Rossia australis] Sarepta [Old Sarepta / Старая Сарепта] / Herb. A. Becker" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p04215442 - P 04215442.jpg - https://www.gbif.org/occurrence/474703068 - https://explore.recolnat.org/occurrence/F75438FF854E4D969C70D1EDF51846C0; rev. Ó. Sánchez Pedraja Jul 2017 [the sheet contains two specimens, only the left-side specimen is P. arenaria]); "E. Boissier . — L. Leresche. — E. Levier / Iter hispanico-Lusitanicum 1878 / Orobanche amethystea Thuill. / Santander ad radices Eryngii amethy maritimi / 15 Jul. legit E. Levier" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p04384533 - P 04384533.jpg - hb. E. Drake - http://www.gbif.org/occurrence/464977980 - https://explore.recolnat.org/occurrence/01FDE30AF2C54581A8E1E721BBE2C880; rev. Ó. Sánchez Pedraja Apr 2013 [the label seems apparently mistaken]); "E. Boissier. — L. Leresche. — E. Levier / Iter hispanico-lusitanicum 1878 / Orobanche amethystea Thuill. / Cantabria: in arena maritima ad Santander, / ... al Saladero [Sardinero (Santander)]; ut videtur, in radic. Eryngii maritimi / 5 Jul. legit E. Levier" (https://data.rbge.org.uk/herb/E01283962 - E 01283962.jpg - hb. Dörfler - https://iiif.rbge.org.uk/herb/iiif/E01283962/full/1600,/0/default.jpg - https://www.gbif.org/occurrence/4420517367; rev. Ó. Sánchez Pedraja Dec 2024, sub P. arenaria)

= Orobanche arenaria f. stenosepala Beck in Biblioth. Bot. 19: 130 (1890). Syn.: ? = Phelipanche atlantica Pomel in Bull. Soc. Sci. Phys. Algérie 11: 103[-104] (1874) [Nouv. Mat. Fl. Atl. 11: 103[-104] (1874 [post 10 Mar 1874])] (Beck 1930: 105). Ind. loc.: "Aus Deutschland [Germany] von Ruinenberg bei Freienwalde (Gallasch), Russland [Russia] und aus dem Kaukasus von Hohenacker gesammelt im Herb. St. Petersburg gesehen" (Beck 1930: 105)

= Kopsia arenaria f. ionantha (A. Kern.) Bég. in Fiori & Paol., Fl. Italia 2: 473 (1902 [Jun 1902]) ["b. ionantha [Kerner sub Orob.]"], nom. rejic. Syn.: ≡ Orobanche ionantha A. Kern. in Oesterr. Bot. Z. 24: 46[236] (1874)

= Orobanche arenaria f. personata Beck in Repert. Spec. Nov. Regni Veg. 18: 35[467] (1922). Syn.: ≡ Phelipanche arenaria var. personata (Beck) Uhlich & Rätzel in Nordic J. Bot. e04155: 5 (2024). Ind. loc.: Ukraine: "Rossia australis: Guv. Jekaterinoslaw [Yekaterinoslav Governorate / Екатеринославская губерния (Dnipropetrovsk Oblast / Дніпропетровська область)] distr. Nowomoskowsk [Novomoskovsk / Новомосковськ] prope Degatowa stanka ad Artemisiae radices, m. Julio 1895 (Alexeenko)" (Beck 1922: 35). Herbarium Specimens: «Lectotype (designated here): A) ‘Rossia australis / Gub. Ecatherinoslaw. Distr. Nowomoskowsk. / Prope Degatowa Stanka legit Alexeenko / [in] Artemisia / 2-14.VII.1895.’ B) ‘Orob[anche] arenaria Borkh. / f. nova personata / Corollae 30–35 mm longae, pronus curvatae / fauce hiante; lacin[iis]. labii inf[eri]. oblongis / acutis mucronatis v[el]. denticulatis / Dr. G. de Beck [m. Beck]’ C) Holotypus O. arenaria f. / personata Beck 1922 / Holger Uhlich, Welterod, Germany (2020)’ (LE 01060138 [http://re.herbariumle.ru/01060138 - LE01060138.jpg], Supporting information)» (Uhlich & Rätzel 2024: 5, sub P. arenaria var. personata)

= Phelypaea arenaria f. ramosa [Merino ex] M. Laínz in Brotéria Ci. Nat. 24(51): 125[18] (1955) ["723 Phelypaea arenaria (Borkh.) Walp. fma. ramosa Merino, f. nov."]. Syn.: ≡ Phelypaea arenaria var. ramosa Merino [before 1906, in sched., nom. nud.]. Ind. loc.: "Caulibus leviter ramosis. Abunde cum typo intermixta variabili satis, ad ostia minniana [Mouth of Miño River, Galicia, Spain / España]". Obs. "Typus formae in herb. Merino (Lourizán), n.º 789 [1526]"

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= Orobanche arenaria subf. calva Beck in Engl., Pflanzenr. 96: 105 (1930). Ind. loc.: Azerbaijan: "Transkaukasien: im Gub. Baku [Baku Governorate / Бакинская губернія] am Kaspischen Meere bei Busovni, im Juni (Petunikow!)"

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Misapplied names:

–  Phelypaea caesia sensu Gren. in Gren. & Godr., Fl. France 2: 624 (1853) ["Ph. caesia Reut. in DC. prodr. 11, p. 6 (non Griseb.)"] [saltem p.p.], non Orobanche caesia Rchb., Iconogr. Bot. Pl. Crit. 7: 48, t. 698 f. 936 (1829). Syn.: = Phelipanche camphorosmae Carlón & al. (2008) (Carlón & al. 2008: 42, MARS); = Phelipanche arenaria (Borkh.) Pomel (1874) ["lèvres ... à dents obtuses" (MA 114850; MA 114851)]; = Phelypaea caerulea (Vill.) C.A. Mey. (1831) [= Phelipanche purpurea (Jacq.) Soják (Coste 1904: 64)]; = Orobanche purpurea Jacq. (1762) [= Phelipanche purpurea (Jacq.) Soják (Tela Botanica - search, accessed Sep 2007)]. Ind. loc.: "Hab. Sur les racines de l'Artemisia gallica Willd. [IPNI] dans les sables de la Mediterranée; Marseille! [France] (Jord.); Banyuls dans les Pyrénées-Or. [France] (Jord.). Ч Juin".

– Kopsia purpurea sensu Samp., Lista Esp. Herb. Portug.: 116 (1913) et Fl. Portug. (2.ª ed.): 499 (1946 [28 May 1947]), non Kopsia purpurea (Jacq.) Bég. in Fiori & Paol., Fl. Italia 2: 473 (1902). Obs. Parasitic on "Artemisia crithmifolia Lin. [= Artemisia campestris subsp. maritima Arcang.] ... Norte" (Sampaio 1947: 499)

– Orobanche purpurea auct. div. [saltem p.p.], non Orobanche purpurea Jacq., Enum. Stirp. Vindob.: 108, 252 (1762)

– Orobanche caerulea auct. div. [saltem p.p.], non Orobanche caerulea Vill., Hist. Pl. Dauphiné 2: 406 (1787) (e.g., Planellas 1852: 325)

– Phelypaea caerulea auct. div. [saltem p.p.], non Phelypaea caerulea (Vill.) C.A. Mey., Verz. Pfl. Casp. Meer.: 104 (1831) (e.g., Colmeiro 1888: 270)

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Ind. loc.: Germany: "Ich fand diese Pflanze häufig bey Arheilgen [Arheilgen, Hessen / Hesse / Hessia] auf eben den dürren Sandhügeln, wo ich auch die vorbefchriebene Planze fand".

Typus / Type: "Type in Berlin [B] ?" (Novopokrovsky & Tzvelev 1958: 53[49]). "Herbarium M[artin]. Dürer. / Orobanche arenaria Borkh. / Flora des Grossherz. Hessen / Fundort Sandfelder bei Arheilgen / unweit Darmstadt. Auf Artemisia camp. / Zeit VI. [18]84 / Leg. M. Dürer / Frankfurt a. M. (Deutschland) FR-110886 (Abb. 1)." ... "Neotypification of Orobanche arenaria Borkh. A specimen in the herbarium Senckenbergianum Frankfurt/Main (FR; fig. 1, 2) collected 1884 by Martin Dürer at Arheilgen near Darmstadt (locus classicus), is chosen as neotype for O. arenaria Borkh." (Uhlich & Gregor 2017).

Designated by: Uhlich & Gregor in Kochia 10: 38 (2017).

Host: mainly on Artemisia species [Compositae] (Annotated Checklist of Host Plants of Orobanchaceae, accessed Jun 2009).

Achillea coarctata Poir. [= Achillea compacta Willd.]. North Macedonia (Teofilovski & Mandzukovski 2023: 97-98, sub P. arenaria; photographies by Z. Nikolov, pers. comm., May 21, 2024, sub P. arenaria).

Artemisia absinthium L. [unconfirmed] (Beck 1930: 103, sub O. arenaria).

Artemisia arenaria DC. [= Artemisia albicerata Krasch.; = Artemisia lercheana Presc. ex Besser]. Russia (http://www.plantarium.ru/page/image/id/515586.html, 515588, 515590, 515591, 515592 by Александр Фатерыга [Alexander Fateryga] 2017; MW 1015091.jpg).

Artemisia campestris L. (Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria), Algeria (Pomel 1876: 103, sub Phelipanche arenaria; P 04076079.jpg - hb. E. Cosson; MPU 261551.jpg - hb. Maire; MPU 261556 - hb. Battandier - Maire), Austria (Kerner 1874: 46, sub O. ionantha), France (Saint-Lager 1883: 394, sub Phelypaea arenaria Walp.; Franchet 1885: 450; MPU - Planchon; P 04384507.jpg - hb. E. Drake), Germany (Dietrich 1835: 151, sub O. robusta; P 04392702.jpg; P 04392701.jpg: Uhlich & Gregor 2017, sub O. arenaria), Poland (Piwowarczyk & al. 2011: 40), Romanie (E 01005562 - hb. Dörfler), Slovakia (WU 0076578), Spain (Montserrat 1957: 349; Alejandre & al. 2004: 41).

Artemisia campestris L. subsp. glutinosa (Gay ex Bess.) Batt. [≡ Artemisia glutinosa J.Gay ex Besser]. Spain (herb. Sánchez Pedraja 12262 et 13687; Sánchez-Gómez & al. 2016: 111, sub O. arenaria).

Artemisia campestris subsp. maritima (DC.) Arcang. [≡ Artemisia campestris var. maritima DC.; = Artemisia crithmifolia L.]. (Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria), Portugal (Sampaio 1913: 116, sub Kopsia purpurea; Sampaio 1947: 499, sub Kopsia purpurea), Spain (MA 114834 - Merino, sub Phelypaea arenaria var. ramosa; Laínz 1955: 125, sub Phelypaea arenaria f. ramosa M. Laínz).

Artemisia chamaemelifolia Vill. Spain (Winkler ex Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria).

Artemisia fragrans Willd. [≡ Seriphidium fragrans (Willd.) Poljakov]. Azerbaijan

Artemisia marschalliana Spreng. [= Artemisia inodora M. Bieb.]. Russia (Becker ex Beck 1890: 131, sub O. arenaria), Russia, Stavropol Krai (https://www.inaturalist.org/observations/120982132 - https://www.gbif.org/occurrence/3858673033 by Шильников Дмитрий Сергеевич [Shilnikov Dmitry Sergeevich]).

Artemisia valentina Lam. [≡ Artemisia herba-alba subsp. valentina (Lam.) Masclans]. Spain (MA 114830).

Artemisia vulgaris L. (Hoehnel ex Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria), Germany.

Comments: Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 1: 6 t. 2, 11-12, 25 (2002, sub O. arenaria); Documentos Jard. Bot. Atlántico (Gijón) 3: 60 f. 3 ["Phylogenetic relationships"], 65 (2005); Documentos Jard. Bot. Atlántico (Gijón) 6: 12 f. 1 [phylogenetic relationships] (2008).

Chromosomatic number: n = 12, 2n = 24 (Měsíček in Měsíček & Jarolímová 1992; Albers & Pusch 1999; Schneeweiss & al. 2004: 441-443, f. 2 (E) "arenaria, 2n = 24 (diakinesis); (F) arenaria (pachytene with DNA fragmentation; see text for details); (G) arenaria, 2n = 24 (AI), arrowheads indicate unseparated bivalents").

Ic.: a, habit (Pontevedra, Spain); b, flowers (ib.); c, opened corolla showing stames and pistil (Isavarre, Spain); d, sub Orobanche arenaria Borckh. [Linnean herbarium (S-LINN) (accessed Feb 2006)].

Distribution: Afghanistan, Algeria, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Czech Republic, Estonia [see P. purpurea subsp. bohemica], France, Georgia, Germany, Hungary, Iran [unconfirmed], Iraq [in error], Israel [in error], Italy, Kazakhstan, Krim, Moldova [unconfirmed], Morocco, North Macedonia, Poland, Portugal, Romanie, Russia, Serbia [unconfirmed], Slovakia, Slovenia [n.v.], Spain, Switzerland, Tunisia [in error], Turkey and Ukraine.

General distribution (Novopokrovskii & Tzvelev 1958: 53[49]); Mediterranean Region distribution; Iberian Peninsula distribution.

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Specimens examined and literature records.

Afghanistan / Afġānistān / Afganistán / Афганистан (Gilli 1979: 303, sub O. purpurea var. tapeina) [unconfirmed]

Baghlan Province: 1. "K.H. Rechinger, Iter Orientale 1965 / Flora of Afghanistan / Orobanche / Prov. Qataghan: Salang Pass, north side. / Stony Juniper slope. / On Artemisia? / 2200 m 1st July, 1965 / Coll. Jennifer Lamond  No. 2230". – 2. "Orobanche purpurea var. purpurea f. purpurea / 1969 det. A. Gilli" (https://data.rbge.org.uk/herb/E01286593 - E 01286593.zip - https://www.gbif.org/occurrence/4420619703; rev. Ó. Sánchez Pedraja May 2024, sub P. arenaria [flowers erect]); "NE: Kataghan [Qataghan Province]: in decl. borealibus jugi Salang, 2200 m, Lamond 2230" (Gilli 1979: 303, sub O. purpurea var. purpurea f. purpurea); "NE: Kataghan [Qataghan Province]: in decl. borealibus jugi Salang, 2200 m, Lamond 2230" (Gilli 1979: 303, sub O. purpurea var. tapeina).

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Algeria / الجزائر‎  / Argelia / Algérie / Algerien (Pomel 1876: 103, sub Phelipanche arenaria; Battandier 1890: 656, sub P. arenaria Walpers)

Batna Province: "Phelipaea arenaria Walp. / Labèse, circle de Batna, province / Constantine, Algérie. / E. Cosson, 14 Juin 1853." (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02986243 - P 02986243.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/10/14/1/P02986243.jpg - https://www.gbif.org/occurrence/438779187 - https://explore.recolnat.org/occurrence/D0BA9062180B4BCDB22CDBD1BF3B627E; rev. Ó. Sánchez Pedraja Jan 2017); "Phelipaea arenaria Walp. / Mídina / Monts Aurès [massif de l'Aurès], province de Constantine, / Algérie / E. Cosson, 10 juin 1853." (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02986243 - P 02986243.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/10/14/1/P02986243.jpg - http://www.gbif.org/occurrence/438779187 - https://explore.recolnat.org/occurrence/D0BA9062180B4BCDB22CDBD1BF3B627E; rev. Ó. Sánchez Pedraja Jan 2017); "Phelipaea arenaria Walp. / ... / ... de Batna, Algérie. / du Colombier, 1854 (n. 253)" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02986243 - P 02986243.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/10/14/1/P02986243.jpg - http://www.gbif.org/occurrence/438779187 - https://explore.recolnat.org/occurrence/D0BA9062180B4BCDB22CDBD1BF3B627E; rev. Ó. Sánchez Pedraja Jan 2017); "Batna" (Pomel 1876: 103, sub Phelipanche arenaria, "sur Artemisia campestris"); "Université d'Alger / Herbier de l'Afrique du Nord / Phelipaea arenaria Walp. / Orobanche / sur Artemisia campestris L.? / C. Aurès, Sgag [forêt de Sgag / forêt Legag?, Aurès Mountains, Saharan Atlas] / corolla bleu violacé / stigmate blanchâtre / 3-7-1920 Dr. R. Maire" (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu261551 - MPU 261551.jpg - hb. Maire - https://www.gbif.org/occurrence/1424916004; rev. L. Carlón & Ó. Sánchez Pedraja 2006-2007 [ut P. arenaria] et Ó. Sánchez Pedraja Nov 2017 [the specimen seems to correspond to Phelipanche purpurea subsp. ballii]); "Batna" (Battandier 1890: 656, sub P. arenaria Walpers, p.p.).

Djelfa Province: "Tadmit [34°30'24.1"N 3°02'13.8"E 34.506687, 3.037164], 5 Jun 2023 / Host Plant: Artemisia campestris" (https://www.inaturalist.org/observations/205785741 - https://www.gbif.org/occurrence/4607438006 by Djoudi Salah Eddine in iNaturalist; accessed May 2024 [!]).

Laghouat Province: 1. "450 / Phelipaea ----- / fl. violettes .-- Sur Artemisia / campestris. / (O.) Sidi Bou Lifâ [Sidi Bou Lefa] (près Aflou) / 5 Juin 88 [1888] / Dr. Clary [J.B.E.Clary]". – 2. "Phelipaea arenaria Walp. / Determinavit: J. A. Battandier / ..." (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu261556 - MPU 261556 - hb. Battandier - Maire - http://www.gbif.org/occurrence/1424916022 - https://explore.recolnat.org/occurrence/34FAF66E2AF1486AA221C8FF90F0219C; rev. Ó. Sánchez Pedraja Nov 2017); "450 / Phelipaea Aegyptiaca Pers. / Sur Artemisia campestris L. ! / Sidi Bou Lifâ [Sidi Bou Lefa, Aflou] / 5 Juin 88 [1888] / Dr. Clary [J.B.E.Clary]" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04076079 - P 04076079.jpg - hb. E. Cosson - http://www.gbif.org/occurrence/474737453 - https://explore.recolnat.org/occurrence/484B93B1F8E7480CA35C1A7986F90BB1; rev. Ó. Sánchez Pedraja Aug 2017); "Aflou [Amour Range, Saharan Atlas]" (Battandier, 1890: 656, sub P. arenaria Walpers).

Oum El Bouaghi Province: "Phelipaea arenaria Walp. / Djebel es Stih Ali prope Batner[?] / 14 Jun. 1869 / A. Letourneux" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02986244 - P 02986244.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/10/14/1/P02986244.jpg - https://explore.recolnat.org/occurrence/806641A4C5B6425185189DD16380B046; rev. Ó. Sánchez Pedraja Jan 2017).

Tiaret Province: 1. "Parabanche caerulea / var. atlantica / itima 25 mai 60 [m. Pomel]". – 2. "... Orobanche laevis L. / = Phelipanche atlantica Pomel! (Type!) / O. Itima, Mou-el-Gtouta [au S. de Tiaret, Hauts plateaux Oranais, Oran department] / 25-5-1860  leg. A. Pomel [m. Maire]" (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu008101 - MPU 008101.jpg - herb. Pomel - http://www.gbif.org/occurrence/1424544992; rev. Ó. Sánchez Pedraja Nov 2009, holotype of Phelipanche atlantica [= Phelipanche purpurea subsp. ballii]); "Sur Artemisia campestris: Mou-el-Gtouta [au S. de Tiaret, Hauts plateaux Oranais, Oran department (see https://dz.geoview.info/zip/20002_el_gtouta)]" (Pomel 1874: 103[-104], loc. class. of Phelipanche atlantica [= Phelipanche purpurea subsp. ballii]).

Tlemcen Province: "Phelipaea arenaria / Aïn Sfissifa [Remchi] / sud de la province d'Oran / Cosson, 5 mar 1856." (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02986242 - P 02986242.jpg - hb. Cosson - http://sonneratphoto.mnhn.fr/2011/10/14/1/P02986242.jpg - http://www.gbif.org/occurrence/439112721 - https://explore.recolnat.org/occurrence/7801DD36C3E94024884CA2AD58E9F3EC; rev. Ó. Sánchez Pedraja Jan 2017 [only the two robust specimens on upper left corner, the other specimens contained in the sheet seem to correspond to P. portoilicitana]).

Unknown locations: "Phelipaea arenaria / sur Artemisia campestris / Teratologie certaines fleurs ant. 8 etaimes / et 2 styles 8 lobes a la corolla" (MPU 261554 - hb. Battandier - Maire; rev. Ó. Sánchez Pedraja Nov 2017 [the sheet contains three specimens, the two upper specimens correspond to P. arenaria (big flowers) and the lower specimen corresponds to P. portoilicitana (smaller flowers)]); idem (MPU 261555 - hb. Battandier - Maire; rev. Ó. Sánchez Pedraja Nov 2017 [the sheet contains two specimens corresponding to P. arenaria (big flowers)).

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Armenia / Hayastan / Armenien / Армения (Koch 1849: 661, loc. class. of Phelypaea arenaria var. elongata; Piwowarczyk & al. 2019: 31-33, f. 20, sub P. arenaria)

Gegharkunik Province / Гехаркуникская область: (Piwowarczyk & al. 2019: 33, sub P. arenaria).

Kotayk Province / Котайкская область: Кавказ, Армения (Армения) [Caucasus, Armenia (Armenia)] "Herbarium Universitatis Mosquensis / M.G. Pimenov. Plantae caucasicae / Orobanche purpurea Jacq. / Армения, ... Гутанасар [Gutanasar] ... / 1997 N.º 1057 / 7791" (https://plant.depo.msu.ru/open/public/item/MW0717054 - MW 0717054.jpg; rev. Ó. Sánchez Pedraja Nov 2017 [the specimens seem to correspond to P. arenaria]) (Piwowarczyk & al. 2019: 33, sub P. arenaria).

No precise location given: "In der Ebene des Araxes [Aras river, Caucasus], c. 2700' hoch" (Koch 1849: 661, loc. class. of Phelypaea arenaria var. elongata).

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Austria / Österreich / Autriche (Kerner 1874: 46, loc. class. of O. ionantha; Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria; Aeschimann & al. 2004: 284, sub O. arenaria; map of P. arenaria in Pflanzenfotos von Stefan Lefnaer [Die Daten in den Tabellen stammen aus Fischer & al., 2008 und wurden von R. Marschner (botanische-spaziergaenge.at) digital aufbereitet.], accessed Jan 2017)

Burgenland / Őrvidék: "Sandige Tro­ckenrasen. Parasitiert auf Feld-Bei­fuß Artemisia campestris und auf Gewöhnlichem Beifuß Artemisia vulgaris. Sehr selten. / Vom Aussterben bedroht!" (photos of P. arenaria in Burgenlandflora, accessed Jun 2017).

Niederösterreich / Lower Austria / Basse-Austriche: "N" (Aeschimann & al. 2004: 284, sub O. arenaria).

Krems-Land: "5.6.2007 ÖK37 76a/59a F131/ 2840 / Phelipanche arenaria / Dürnstein - Krems (Weingartenbereich)" (http://www.botanische-spaziergaenge.at/viewtopic.php?f=86&t=1923 in botanische-spaziergaenge.at, accessed Feb 2014); "3601 Dürnstein [48°23'34.9"N 15°31'54.0"E 48.393017, 15.531668], 8 Jun 2022" (https://www.gbif.org/occurrence/3859945008 - https://www.inaturalist.org/observations/120866556 by https://www.inaturalist.org/people/auguste71 [Ingrid Schmitzberger] in iNaturalist; accessed May 2024 [only the photo https://inaturalist-open-data.s3.amazonaws.com/photos/204473204/original.jpeg [P. arenaria] the rest of photos correspond to P. purpurea subsp. bohemica [https://inaturalist-open-data.s3.amazonaws.com/photos/204470818/original.jpeg, .../204473192/original.jpeg, .../204473212/original.jpeg and ...204473235/original.jpeg], all apparently on Artemisia sp.]).

Salzburg / Såizburg / Salzburgo: "S" (Aeschimann & al. 2004: 284, sub O. arenaria).

Tyrol / Tirolo: 1. "Orobanche ionantha K. / Tirol. Trautson bei Matrei [Matrei in Osttirol (Lienz District)] / Auf Artemisia campestris / Kerner". – 2. Syntypus WU 0076573 / Orobanche ionantha A. Kern. / ... / typifcavit M. Lachmayer (WU) 2014-05-21 / ..." (https://wu.jacq.org/WU0076573 - https://www.jacq.org/detail.php?ID=482238 - WU 0076573.jpg - https://object.jacq.org/europeana/WU/482238.jpg - https://www.gbif.org/occurrence/1830990375; rev. Ó. Sánchez Pedraja May 2024, syntypes of O. ionantha); "Unter den in Ungarn [Hungary] gesammelten Orobanchen fand sich auch ein Exemplar einer noch nicht beschriebenen Art, welche ich seither auch an mehreren Punkten in den Thälern der Alpen auf Artemisia campestris auffand, und die ich insbesonders in der Nähe von Innsbruck [Tyrol] auf den Angern der sogenannten "Höttinger Aue [Höttinger Au]"..." (Kerner 1874: 46, loc. class. of O. ionantha); "Dr. C. Baenitz, Herbarium Europaeum Nº ... / Orobanche ionantha A. Kern. / O. cerulea Vill proparte / Tirol: Ad radices Artem. camp. circa / Innsbruck, raro 1882.VI / leg. J. Murr." (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04419637 - P 04419637.jpg - http://www.gbif.org/occurrence/465125215 - https://explore.recolnat.org/occurrence/779BED04FAD54B019FDCBF765B6420AA; rev. Ó. Sánchez Pedraja May 2018); idem (LY 0737615.jpg - hb. Rouy in Roland Bonaparte - https://www.gbif.org/occurrence/3427999059 - https://explore.recolnat.org/occurrence/C44CE22AE4804B30A725E257ADD29951; rev. Ó. Sánchez Pedraja May 2024); "Tirol: am Ufer des Im bei Imst / auf Artemisia camp. ... / Aug. 1902 / Leg. Louis Keller" (MPU - Faure, sub O. purpurea, rev. L. Carlón & Ó. Sánchez Pedraja 2006-2007); "NT" (Aeschimann & al. 2004: 284, sub O. arenaria).

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Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaïdjan [former Azerbaijan Soviet Socialist Republic [Azerbaijan SSR] / Азербайджанская Советская Социалистическая Республика [Азербайджанская CCP] (Beck 1930: 104, sub O. arenaria; Beck 1930: 105, loc. class. of O. arenaria sub f. calva; Karjagin 1957)

Absheron-Khizi Economic Region / Abşeron-Xızı iqtisadi rayonu: "Plants from the Southern Caucasus / ... / ... / 13629. Orobanche cf. purpurea / Azerbaijan.– Xixi rayonu [Khizi District / Xızı rayonu]: Abşeron Yarımadesı [Absheron peninsula], / Tugcay valley ca 5-8 km SW Giləsi [Giləzi / Gilazi] along the Giləsi - Xizi / highway,  ca. 50-80 m. / Marly slopes supporting Artemisia frangrans steppe / Coll. Thomas Borsch, Vüqar Kǝrimov, Nadja Korotkova / & Gerald Parolly [Pa 13621-13629a] / Det.: G. Parolly / 25.04.2013". – 2. "Phelipanche cf. iberica (Beck) Tzvel. / rev.: Holger Uhlich ... 2014". – 3. "Phelipanche iberica (Beck)  Tzvel. ... / rev. H. Uhlich & S. Rätzel / 9.02.2022" (B 10 0449859 - https://herbarium.bgbm.org/object/B100449859 - https://www.jacq.org/detail.php?ID=1802628 - https://www.gbif.org/occurrence/3763006337; rev. Ó. Sánchez Pedraja Aug 2022, sub P. arenaria [compared to https://www.inaturalist.org/observations/48921984]); "Plants from the Southern Caucasus / ... / ... / 13629. Orobanche cf. purpurea / Azerbaijan.– Xixi rayonu [Khizi District / Xızı rayonu]: Abşeron Yarımadesı [Absheron peninsula], / Tugcay valley ca 5-8 km SW Giləsi [Giləzi / Gilazi] along the Giləsi - Xizi / highway,  ca. 50-80 m. / Marly slopes supporting Artemisia frangrans steppe / Coll. Thomas Borsch, Vüqar Kǝrimov, Nadja Korotkova / & Gerald Parolly [Pa 13621-13629a] / Det.: G. Parolly / 25.04.2013" (B 10 1259349.jpg - http://herbarium.bgbm.org/object/B101259349 - https://www.jacq.org/detail.php?ID=2076971 - https://www.gbif.org/occurrence/5029332555; rev. Ó. Sánchez Pedraja Feb 2025, sub P. arenaria).

Mountainous Shirvan Economic Region / Dağlıq Şirvan iqtisadi rayonu: "im Gubern. Baku [Baku Governorate / Бакинская губернія], Distr. Schemacha [Shamakhi / Şamaxı] bei Dghenga, im Mai (Petunnikow!)" (Beck 1930: 104, sub O. arenaria); "Transkaukasien: im Gub. Baku [Baku Governorate / Бакинская губернія] am Kaspischen Meere bei Busovni, im Juni (Petunikow!)" (Beck 1930: 105, loc. class. of O. arenaria sub f. calva).

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Belgium / Belgique / Belgien / Bélgica (Franchet 1885: 450, Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria)

Flemish Region / Vlaams Gewest: "Flandria: Inter Nieuport [Nieuwpoort] et Coxyde [Koksijde]" (Beck 1890: 131, sub O. arenaria).

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Bulgaria / България / Bulgarie / Bulgarien (Velenovský 1891: 439, sub P. arenaria; Velenovský 1898: 222, loc. class. of Phelypaea arenaria var. euxina; Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria; Stoyanov 2005: 462-463 f. 1 [distribution map], sub O. arenaria; Stoyanov & al. 2015, sub P. arenaria)

Dobrich Province / Област Добрич: "Ad Balčik [Balchik / Балчик], Kavarna [Каварна] ad Pontum [Pontus Euxinus / Black Sea (Varna)] ‭(Str) ..." (Velenovský 1898: 222, sub Phelipaea arenaria).

Pazardzhik Province / Област Пазарджик: "34TGM45 (17.1) in collis saxosis mt. Rhodope occidentalis [Rhodope Mountains] Cepino [Chepino Valley / Чепинска котловина / Chepino / Чепино], 811 m, 23.07.1925, SV (Davidov), SOM 69762; Cepino [Chepino Valley / Чепинска котловина / Chepino / Чепино] ad pagum Lazene, 750 m, 24.06.1926, SV (Davidov), SOM 69763, SOM 69768 (sub O. purpurea); 35TKG85 (17.1), over Bratzigovo, 440 m, parasite on Achillea, 7.08.2003, SC (KS), SOA s/n" (Stoyanov 2005: 462, sub O. arenaria).

Plovdiv Province / Област Пловдив: "Bulgarien: bei Kara afly [Plovdiv Province] (Pichler!)" (Beck 1930: 103, sub O. arenaria); "35TLG15 (17.2) over Markovo, 400 m, parasite on Achillea pannonica, 1.06.2003, SC(KS), SOA 56536" (Stoyanov 2005: 462, sub O. arenaria).

Sofia Province / Софийска област: "Ph. arenaria Walp. (Ph. purpurea Jcq.). In declivibus m. Balkan versus Sofiam [Sofia / София] (Vel)" (Velenovský 1891: 439, sub Ph. arenaria [see Phelipanche purpurea]).

Varna Province / Област Варна: 1. "Flora bulgarica / Phelypaea arenaria Walp. ! Beck [m. Beck] / β) euxina m. Varna (Stribrny) / 1897.V Legit [det.] dr. J. Velenovský". – 2. "Orobanche arenaria Borkh. / 1970 Revidit: K. Kult". – 3. "Typus" (PRC 451846 - https://prc.jacq.org/PRC451846 - https://www.jacq.org/detail.php?ID=559522 - https://www.gbif.org/occurrence/3413676613 - http://www.jacq.org/image.php?filename=559522&method=europeana; rev. Ó. Sánchez Pedraja Jul 2022, type of Phelypaea arenaria var. euxina); "... Varna ‭(Sk)" (Velenovský 1898: 222, sub Phelipaea arenaria); "Pontum [Pontus Euxinus / Black Sea (Varna)] ‭(Str) ..." (Velenovský 1898: 222, sub Phelipaea arenaria); "Varna (Stribrny)" (Beck 1930: 103, sub O. arenaria).

Other locations: "Bulgaria: prope Tulscha [Tulcea] Dobrutschae [Dobruja, Romanie] leg. Sintenis" (Beck 1890: 131, sub O. arenaria); "bei Tulscha [Tulcea] in der Dobrudscha (Sintenis!). — Bulgarien: bei Kara afly [Plovdiv Province] (Pichler!), Varna (Stribrny)" (Beck 1930: 103, sub O. arenaria).

"New and unpublished data" (Stoyanov 2005: 462-463 f. 1 [distribution map], sub O. arenaria).

Note: The host plants of O. arenaria are, according to the herbarium data (Table 2), Achillea - A. millefolium L., A. coarctata Poir. (FB4; FRB), A. pannonica and Stachys recta L. The host plants Artemisia (FB4; KBP; KPB3) - A. campestris (FB1), Anthemis (FB4) and other Asteraceae (FRB; KPB3), Eryngium maritimum L. (FB4), Lamiaceae (KVPB) are indicated, but this information is not confirmed by herbarium data." (Stoyanov 2005: 463, sub O. arenaria); "Гостоприемници от флората на България: Apiaceae: Eryngium maritimum, Asteraceae: Achillea coarctata, A. millefolium, A. nobilis, A. pannonica, Artemisia sp. div., Tanacetum millefolium; Lamiaceae: Stachys recta" (Stoyanov & al. 2015, sub P. arenaria).

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Croatia / Hrvatska Croacia / Kroatien / Croatie (Nikolić 2000; Nikolić 2015, sub O. arenaria)

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Czech Republic / República Checa / Česká republika (Beck 1930: 103, sub O. arenaria; Zázvorka 2000: 484-485, t. 78 f. 4!, sub P. arenaria)

South Moravian Region / Jihomoravský kraj

Břeclav District: "7.6.2007 ÖK11 71b/65b F131/ 2840/ Phelipanche arenaria / Mikulov / Nikolsburg" (http://www.botanische-spaziergaenge.at/viewtopic.php?f=86&t=1923 in botanische-spaziergaenge.at, accessed Feb 2014).

Znojmo District: Ječmeniště [Vrbovec] (https://www.biolib.cz/en/image/id223099 by Jiří Kameníček in BioLib.cz, accessed May 2018).

Ústí nad Labem Region / Ústecký kraj

Teplice Distric: České středohoří Mountains, Štěpánovská hora Hill 28th June 2009 (photo no. 12169b by Ivan Bilek in Naturephoto-cz, accessed Feb 2014).

Louny District : Podbořany [Podersam] (https://www.biolib.cz/IMG/GAL/BIG/155670.jpg by Daniel Koutecký in BioLib.cz, accessed May 2018).

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Estonia / Estland / Eesti / Эстони [see P. purpurea subsp. bohemica]

Võru maakond / Võru County: 1. "VII.1893 / Neu Kasseriz (= Kasaritsa) bei Werro (= Võru lähedal) / auf Artemisia campestris / leg. H. Hiir". – 2. "Orobanche arenaria Borkh. / Sandsommerwurz". – 3. "Orobanche purpurea Jacq. / L. Viljasoo / 10.XI-65." (TU 301922 [Tiina Samson]; rev. Ó. Sánchez Pedraja Dec 2017 [it seems to correspond to P. purpurea subsp. bohemica parasitic on Artemisia campestris]).

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France / Francia / Frankreich (Cosson & Germain 1840: 63, sub O. arenaria Borkh.; Reuter 1847: 6, sub Phelipaea arenaria; Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria; Tela Botanica, accessed Jan 2010; Faune Flore Fonge Massif Central, accessed Jan 2017; Floraphile 45 taxon 850, accessed Jul 2017)

Auvergne-Rhône-Alpes [Auvergne-Rhône-Alpes]

Haute-Savoie (74): "1825 et 1826, sur les collines qui bordent la route de Saint-Pierre à Sion [Rhône-Alpes], à demi-lieue de cette dernière ville. Elle est exclusivement parasite de l’Artémise vulgaire, comme l’indique aussi Wallroth…". Obs. "anthères … cotonneuses" (Vaucher, 1827: 65, loc. class. of Orobanche de l'Artemise commune); "Chamonix" (Saint-Lager 1883: 394, sub Phelypaea arenaria Walp.).

Haute-Loire (43): Le-Puy-en-Velay, «sur l’Artemisia campestris», 20-VII, J.E. Planchon (MPU - Planchon; rev. Carlón & Ó. Sánchez Pedraja 2006-2007 [the sheet consists of five specimens, the three lower ones seem to be Phelipanche inexspectata (1 left) and P. arenaria (2 right) (Carlón & al. 2008: 33). The rest are Orobanche caryophyllacea from Ostende dunes (Belgium)] (Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria).

Isère (38): Mizoën, 19.06.2004, sobre Artemisia campestris L. (?), pastos pedregosos, secos y soleados, G. Gómez Casares & G. Moreno Moral (obs.).

Puy-de-Dôme (67): "Ch. d'Alleizette / plantes d'auvergne / Phelipaea arenaria / Sur Achillea millefolium / Puy-de-Dôme. Puy Chaleix, pelouses. / 20-VI-1932" (https://science.mnhn.fr/institution/clf/collection/clf/item/clf170369 - CLF 170369.jpg - https://www.gbif.org/occurrence/1437901222 - https://explore.recolnat.org/occurrence/CBAA41F6026D40B08FAFF63D0385DBEA; rev. Ó. Sánchez Pedraja Jul 2017 [all specimens correspond to P. purpurea apparently parasitizing Achillea sp.]).

Rhône (69D): "Parasite sur les racines de l'Artemisia campestris ... la Pape" (Saint-Lager 1883: 394, sub Phelypaea arenaria Walp.).

Centre-Val de Loire

Eure-et-Loir (28): "Néron" (Saint-Lager 1883: 394, sub Phelypaea arenaria Walp.).

Indre-et-Loire (37): "Herbier Bernard Guémas // Nº 3330 AR Herbier 753 / ... / Orobanche amethystea Th. / ... / 3 Juin 1989 ... / ... / coteaux ... (sur Eryngium campestris [the host is mistaken?]) / sablonneux / Candes [Candes-Saint-Martin] (vienne [Vienne]) / ..." (ANG 003107; rev. Ó. Sánchez Pedraja Nov 2018); "Candes-Saint-Martin [47°12'45.6"N 0°04'52.4"E]" 21 Jun 2014 (https://www.inaturalist.org/observations/52846880 by Susan Waltert, sub P. purpurea ssp. bohemica, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jun 2021 [calyx-teeth are very long and stigma is white, cf. P. arenaria]).

Loir-et-Cher (41): "Phelypaea arenaria Walp. / Soings [Soings-en-Sologne], routin d'une vigne qui / descend a pic au R. du lac. / (Loir-et-Cher) / Juin 1874 / Sur l'Artemisia Campestris / A. Franchet" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04384507 - P 04384507.jpg - hb. E. Drake - http://www.gbif.org/occurrence/464878845 - https://explore.recolnat.org/occurrence/9E21599ED95D47FC88F5BD14444A9FBF; rev. Ó. Sánchez Pedraja Nov 2011); "parasite sur l'Artemisia Campestris. R. Pentes du coteau dominant le lac de Soings au N. et au N.-O.!; sables de la Loire à Nouan [Nouan-le-Fuzelier]! (Lefrou, roger), à Suèvres [Blois], à Rilly [Rilly-sur-Loire, Blois] (Delaunay) surtout dans les iles du fleuve" (Franchet 1885: 450).

Île-de-France

Essonne (91): "Orobanche arenaria Borck. / Lardy / ... Juin 1846. / Herb. Maire" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p04404585 - P 04404585.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/12/29/6/P04404585.jpg - http://www.gbif.org/occurrence/465116014 - https://explore.recolnat.org/occurrence/6BD5594D83C34836BE28CC4209057F4E; rev. Ó. Sánchez Pedraja Nov 2011; "2.942 / Herbier G. Didier / Phelipoea arenaria Walp. / Anthères velues laineuses / Seine & Oise: Sablière de Lardy.- / 26 Juillet 1925 G.D." (P 03931912; rev. Ó. Sánchez Pedraja Jul 2017).

Seine-et-Marne (77): "Nemours (Devilliers)." (Cosson & Germain 1840: 63, sub O. arenaria Borkh.); "Phelypaea arenaria Walp. / Orobanche arenaria Borkh. / Coteaux greseux - calcaires à Nemours, / parasite sur l'Artemisia campestris. / 30 juin 1881. / Herb. Maille" (P 04404582 - hb. E. Cosson; rev. Ó. Sánchez Pedraja Nov 2011); "Orobanche comosa Wallroth / Orobanche arenaria Baechmann. / Roches da Chantriaville / près Nemours" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p04404585 - P 04404585.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/12/29/6/P04404585.jpg - http://www.gbif.org/occurrence/465116014 - https://explore.recolnat.org/occurrence/6BD5594D83C34836BE28CC4209057F4E; rev. Ó. Sánchez Pedraja Nov 2011); "Herbier E. Jeanpert / Phelypaea arenaria Walp. / Nauteau [Nanteau-sur-Essonne] près Malesherbes / champs sableux sur / Art. campestris / 6.7.98 / (Seine-et-Marne)" (P 03996073; rev. Ó. Sánchez Pedraja Oct 2016); "Phelypaea arenaria Walp. / sur Artemisia campestris. / Mt. de Rubrett près / La Grande-Paroisse (S et M) / endroit sablonneux / juillet 1913 / ..." (P 04404584; rev. Ó. Sánchez Pedraja Jun 2017 [the sheet contains five specimens, the four specimens on the left-side (Chamarande) are P. purpurea and the other specimen (La Grande-Paroisse) on the right-side is P. arenaria]); "Ch. d'Alleizette / Phelipaea / Sur Achillea millefolium / Seine-et-Marne - Env. de Nemours / près du canal / sol calcaire / 28 Juin 1929" (http://coldb.mnhn.fr/catalognumber/clf/clf/clf170370 - CLF 170370.jpg - https://www.gbif.org/occurrence/1437901253 - https://explore.recolnat.org/occurrence/C9336F324A214CEC911F0EEBCA3DC325; rev. Ó. Sánchez Pedraja Jul 2017, sub P. purpurea).

Occitanie / Occitània [Languedoc-Roussillon-Midi-Pyrénées]

Aude (11): "Leucate, graviers de la Tet vers St-Estève" (Saint-Lager 1883: 394, sub Phelypaea arenaria Walp. [see O. cernua]).

Gard (30): "Orobanche lavandulacea Reichenbach / Les dunes au nord d'Aigues-Mortes 6 juin 1843 [Foudras Eugène]" (LYJB 006229 - http://mediaphoto.mnhn.fr/media/1531414951707inoAWA90znGXuBjM - https://explore.recolnat.org/occurrence/5F01A80C458B47D3815CD700C3F1F682 - https://www.gbif.org/occurrence/3415123459; rev. Ó. Sánchez Pedraja Oct 2022, sub P. arenaria); "2808 / Herbier J.M. Rouet / Phelypaea purpurea Asch. / (sur Artemisia) / Gard: / la Capelude [la Capedule. (Camargue-ouest)] / 24.5.1965" (P 00598732 - hb. Jean Guillet; rev. Ó. Sánchez Pedraja Nov 2011 [see P. purpurea subsp. bohemica]); "Phelypaea arenaria Borck. / Bois de Broussan [près de Nîmes] Gard / Sur l'artemisia campestris / N." (P 04419486 - Herbier donné par M. Loret; rev. Ó. Sánchez Pedraja Nov 2011); 1. "Herbarium Vadense / ... / Orobanche cf. fuliginosa Reut. / Det.: A. Touw. / Leg.:. A. Touw. No: 612 / Die: 9-6-1959. / Loc.: Zd, Frankrijk. Duinen tussen Carnon en le Grau du Roi [Le Grau-du-Roi]". – 2., "612 / ... / Orobanche cf. fuliginosa Reut. / als vorige; op Athemis / maritima [?] / 9 VI 1959" (WAG 1141278; rev. Ó. Sánchez Pedraja Sep 2016).

Haute-Garonne (31): "H. Sur les racines de l’Artemisia campestris. C. Toulouse [N 43° 36' 15'' E 1° 26' 37'']: à Pech-David, à Venerque sur les côteaux R. R.; les rives sableuses de l’Ariége et de la Garonne. C. C." (Noulet, 1837: 489).

Hérault (34): "Habit Monspelii [Montpellier]" (Linnaeus 1753: 632, loc. class. of Orobanche laevis L. (1753), nom. rej.); "Montpellier" (Grenier 1853: 625); "Orobanche amethystea Thuill. / ... / Plateau volcanique [Roquehaute / Roque-Haute, Portiragnes (Béziers)] / May 1870 / B. Biche" (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu433515 - MPU 433515.jpg - hb. P.B. Biche - https://explore.recolnat.org/occurrence/8F483E1B071844459C94E7861A20B9B2; rev. Ó. Sánchez Pedraja Dec 2024, sub P. arenaria [Flore des environs de Roquehaute par M Biche officier]); "L. Chevalier, Précigné / Société dauphinoise, 1886. / Nº 4992 Phelipaea arenaria Walp. ... / ... / La Motte, près de Montpellier (Hérault): sables / maritimes, sur l'Artemisia gallica Willd. / 2 juin 1885. / Ch. Dupin" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p03996074 - P 03996074.jpg - http://sonneratphoto.mnhn.fr/2012/03/21/1/P03996074.jpg - http://www.gbif.org/occurrence/607793604 - https://explore.recolnat.org/occurrence/55C32B932302492BA4E371FAF5538BEB; rev. Ó. Sánchez Pedraja Oct 2016); "CH. D'Alleizette / Phelipaea arenaria Walp. / Hérault: La Motte [La Grande-Motte], près Montpellier / sables sur Achillea setacea [Achillea millefolium subsp. setacea (Waldst. & Kit.) Čelak.] / 2 Juin 1912" (https://science.mnhn.fr/institution/clf/collection/clf/item/clf170371 - CLF 170371.jpg - http://www.gbif.org/occurrence/1437901227 - https://explore.recolnat.org/occurrence/E033F4D3D7E7443FB87530DAA9FD8957; rev. Ó. Sánchez Pedraja Jul 2017 [the sheet contains three specimens, only the central specimen that is attached to its host (Achillea) is P. purpurea; the two lateral specimens are P. arenaria (very typical and well preserved the left-hand specimen)]); 1. "Orob. arenaria / V (Vendres?) J 90 / ex herb. F. Mouret". – 2. "Orobanche purpurea Jacq. / 24.01.3003 / Michaux & J. Molina ..." (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu993854 - MPU 993854.jpg - hb. Albaille - https://explore.recolnat.org/occurrence/2FA2B81B464E42A6A4E3105BDEDA12DE; rev. Ó. Sánchez Pedraja Dec 2024, sub P. arenaria).

Pyrénées-Orientales / Eastern Pyrenees / Pirineus Orientals / Pirenèus Orientals / Pirineos Orientales (66): "Banyuls-sur-Mer, sur le racines de l'Artemisia Gallica" (Companyo 1864: 515, sub P. caesia); "graviers de la Tet [Têt] vers St-Estève [Saint-Estève / Sant Esteve del Monestir]" (Saint-Lager 1883: 394, sub Phelypaea arenaria Walp.); "Pyr. Atax. ad Salelles [Saleilles, N 42° 39' 0'' E 2° 57' 0''], die 15. Jun 1839; in Pyr. or. sub Prats de Mollo, au Bauze de l'Aze [Cambras d'Azé N 42° 27' 0'' E 2° 8' 0''], die 26. Jun. 1840" (Bubani 1897: 258); "...61 / Herbier E - J - Neyraut. / Phelypaea arenaria Walp. ... / 8 Juin 1890. Pyrénées-Orientales: — Anciens graviers de la Tet [Têt], vis- / à-vis Saint Estève près de Perpignan. // Sur les racines de l'Artemisia campestris" (P 04392355; rev. Ó. Sánchez Pedraja Dec 2015); "Trencada d’Ambouilla, riv. gauche, 15-V-1897, Fre. Sennen, MPU s/n-Herbier Coste" (Pujadas, 2001: 74, sub Orobanche arenaria); "Pentes de Caurinya vers Fillols, 23-VI-1898, Fre. Sennen, MPU s/n-Herbier Coste" (Pujadas 2001: 74, sub O. arenaria).

Provence-Alpes-Côte d'Azur

Alpes-Maritimes (06): "2808 / Phelypaea caerulea / C.A. Mey. / Alpes maritimes - St Martin-Vésubie [Saint-Martin-Vésubie] / ...long de la route de Venanson / 24 juin 1944" (P 00598731 - hb. Jean Guillet, rev. Ó. Sánchez Pedraja Nov 2011); Arrondissement de Nice, "Val. Roya: sur les Artemisia au-dessus de la ville de Tende [N 44° 5' 0'' E 7° 36' 0''], 24.7.1879. Préalp. Menton: au-dessus du village de Peille, talus herbeux, calc., 600 m, 3.6.1903. Val. Tinée: entre Pont-Haut et St-Dalmas le Selvage [Saint-Dalmas-le-Selvage, N 44° 17' 0'' E 6° 52' 0''], rocailles, cristallim, 1300 m, sur Artemisia campestris, 29.7.1905; id., 10.7.1908" (Charpin & Salanon, 1985: 253, sub O. arenaria); "Herbier P. Litzler / Orobanche purpurea Jacq. / France (Alpes-Maritimes): St-Mart.-Vésubie [Saint-Martin-Vésubie] / Belvédère, les Blancons / sur Artemisia campestris / 10.07.1988 / leg. ipse  nº 88/776" (MPU 339612, rev. Ó. Sánchez Pedraja June 2017).

Bouches-du-Rhône (13): "Phelipaea caesia Reut. / Fos [Fos-sur-Mer]: le long du Canal de navigation sur les / racines des Artemisia glutinosa / Mai 1892 / Autheman" (AIX 027159, rev. Ó. Sánchez Pedraja Jun 2017).

Var (83): "Phelipaea caesia Reut. / Sur l'Artémise, au quartier de l'Aréna / Roquebrune [Roquebrune-sur-Argens] (Var) 5 Juin 1899 [m. F.C. Bertrand] / F.C. Bertrand / Instituteur / Roquebrune (Var) [o.s.]" (MA 114850, rev. Ó. Sánchez Pedraja 11 Nov 2007); "4777.- Phelipaea caesia Reut. ... /  Var: Roquebrune, terrains sablonneux à l'Arena / Legit Bertrand / 1er juin 1900  Albert" (MA 114851, rev. Ó. Sánchez Pedraja 11 Nov 2007); idem (AIX 002057, rev. Ó. Sánchez Pedraja Feb 2014); idem (https://science.mnhn.fr/institution/clf/collection/clf/item/clf210005 - CLF 210005.jpg - https://www.gbif.org/occurrence/1437940014 - https://explore.recolnat.org/occurrence/E07E6F2B014E4510B39581BC65ABBC52; rev. Ó. Sánchez Pedraja Jun 2017); "sables maritimes à St. Raphaël (Var), 3.6.1873, leg. Huet sub nom. O. cernua Loefl. [sur Artemisia]" (Charpin & Salanon, 1985: 253, sub O. arenaria).

Vaucluse (84): Départ. du Vaucluse, campo militar de Orange (pr. Orange), 02.06.2006, aparentemente sobre Artemisia campestris L. (Compositae), G. Gómez Casares & G. Moreno Moral MM0120/2006 (herb. Sánchez Pedraja 12592).

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Georgia / Sakartvelo / Georgien / Géorgie / Грузия (former Georgian Soviet Socialist Republic [Georgian SSR] / Грузинская Советская Социалистическая Республика [Грузинская ССР]) (Boissier 1879: 496, Phelipaea arenaria; Beck 1930: 104, sub O. arenaria; Gagnidze 1999: 248, sub O. arenaria)

Kakheti / Кахетия: "Sagarejo, Kakheti [41°26'51.0"N 45°22'32.9"E - 41.447506, 45.375816, near Davit Gareji Lavra Orthodox church (Akhaltsikhe Municipality)], 8 Jun 2019" (https://www.inaturalist.org/observations/28047963 - https://www.gbif.org/occurrence/2283118220 by Felix Riegel in iNaturalist; accessed Jul 2022 [apparently on Artemisia sp.]).

Mtskheta-Mtianeti / Мцхета-Мтианети: "in Ossetien im Engpasse Gudamakar [Gudamaqari] oder H. Aragwa [Aragvi] (Londis!)" (Beck 1930: 104, sub O. arenaria).

Samtskhe-Javakheti Region / Самцхе-Джавахети: "Akhaltsikhe, Samtskhe-Javakheti [41°36'35.6"N 43°05'24.2"E 41.609889, 43.090056, near Rustavi], 16 Jun 2019" (https://www.inaturalist.org/observations/29792504 - https://www.gbif.org/occurrence/2366029838 by Felix Riegel in iNaturalist; accessed Jun 2021 [apparently on Artemisia sp.]).

Tbilisi / Tiflis / Тбилиси: "in Iberiâ prope Tiflis [Tbilisi] (ex cl. Rchb. fil.)" (Boissier 1879: 496, Phelipaea arenaria); "Tiflis [Tbilisi]! (nescio quis legerit)" (Reichenbach f. 1862: 87, sub P. arenaria Walp.).

No precise location given: "Aus Grusien von K. Schmidt" (Koch 1849: 661, sub P. arenaria Borkh.); "Transkaukasien: in Georgien (Wilhelms, Hohenacker!)" (Beck 1930: 104, sub O. arenaria).

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Germany / Deutschland / Alemania (Borkhausen 1822: 17, loc. class. of Orobanche arenaria; Wallroth 1822: 314, loc. class. of Orobanche comosa Wallr.; Walpers 1844: 460, loc. class. of Orobanche robusta A. Dietr.; Reuter 1847: 9, sub Phelypaea robusta; Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria)

Baden-Württemberg (DE1): "Phelypaea arenaria Walp. / 1870 25 Juni Mühlenbach / Siibenbürgen / ... auf Artemisia / ... / Johann v. Csató" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p04384518 - P 04384518.jpg - hb. E. Cosson - http://www.gbif.org/occurrence/465113322 - https://explore.recolnat.org/occurrence/5EF95698209644A5A5981C1ABE54410A; rev. Ó. Sánchez Pedraja Jan 2017); "F. Schultz, herbarium normale, nov. ser. Cent. 4. / 313. Orobanche arenaria Borkh. in Roem. ... p. 625 / 30. julii 1872. / In arenosis prope S. Ilgen (Heidelber, Baden) Solo / siluv. / Leg. Dr. J. Scriba" ( http://coldb.mnhn.fr/catalognumber/mnhn/p/p04385328 - P 04385328.jpg - hb. E. Drake - http://sonneratphoto.mnhn.fr/2011/12/28/2/P04385328.jpg - http://www.gbif.org/occurrence/464720013 - https://explore.recolnat.org/occurrence/7E2AB7124B39427496BB9C2E1E3A62DB; rev. Ó. Sánchez Pedraja Nov 2011); idem (http://coldb.mnhn.fr/catalognumber/mnhn/p/p04393738 - P 04393738.jpg - hb. F.W. Schultz - http://sonneratphoto.mnhn.fr/2011/12/30/4/P04393738.jpg - http://www.gbif.org/occurrence/465178112 - https://explore.recolnat.org/occurrence/9386A2C6E0814945A354E88352439DE5; rev. Ó. Sánchez Pedraja Jan 2017); idem (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04384527 - P 04384527.jpg - hb. E. Cosson - http://www.gbif.org/occurrence/464964064 - https://explore.recolnat.org/occurrence/45C24AB152E544B59EA952376327858F; rev. Ó. Sánchez Pedraja Jun 2017 [the sheet contains five specimens, only the three specimens on the right-side of the sheet are P. arenaria (near their label), the other two are P. purpurea]).

Brandenburg (DE4): "Orobanche arenaria / Borkh. / collegi pr. Frankfurt ad Viadr [= Oder] / parasitice Artemisia campestris / J.N. Buek" (P 04392702; rev. Ó. Sánchez Pedraja Jul 2017); "Nº 723 Phelypaea arenaria Walp. [...] Près de Lychen (Brandenburg, Prusse: sur Artemisia / campestris / Juillet 1890 W. Bertram) (MA 114837, rev. Ó. Sánchez Pedraja 11 Nov 2007); "Crescit in radicibus Artemisiae campestris in collibus arenosis apricis (bei Baumgartenbrück inter Potsdam)" (Walpers 1844: 460, loc. class. of Orobanche robusta A. Dietr.); "Auf sonnigen Anhöhen im Sandboden bei Baumgartenbrück inter Potsdam im Jarhr 1834 in Gemeinschaft mit dem Herrn Cabinets … und in Büschen wahrscheinlich auf Artemisia campestris wächst. Blüht im Juli" (Reuter 1847: 9, sub Phelypaea robusta, "Borussiae prope Potsdam [loc. class.]"); "Herb. Acad. Techn. Delph-Bat. / Orobanche arenaria Borkh. / Prenzlau: Hindenburg [Hindenburg near Prenzlau]. 1075 / Coll. Swart ontv. Mei 1930 / leg.: Grantzow" (https://data.biodiversitydata.nl/naturalis/specimen/L.2817855 - L 2817855.jpg; rev. Ó. Sánchez Pedraja Jun 2021 [sub P. purpurea subsp. bohemica]).

Bayern / Baviera (DE2): Bavaria: München (Beck 1890: 131, sub O. arenaria).

Hessen / Hesse (DE7): "Ich fand diese Pflanze häufig bey Arheilgen auf eben den dürren Sandhügeln..." (Borkhausen 1822: 17, loc. class. of Orobanche arenaria); "Flora Galliae et Germaniae exsiccata / 495. Orobanche arenaria, Borkhausen ... / ... O. comosa, Wallr. / 16 Juillet 1840 / Sur les racines de l'Artemisia campestris, dans les sables in- / cultes, près de Darmstadt (Hesse); rec. G. Gravelius." (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04392701 - P 04392701.jpg - http://www.gbif.org/occurrence/464720008 - https://science.mnhn.fr/images/recolnat.png; rev. Ó. Sánchez Pedraja Jul 2017); "Orobanche arenaria, Borkh. / Darmstadt / Coll. Gravelius, 1843" (US 147296 - Barcode 03926920 - http://n2t.net/ark:/65665/3b91c2d15-e238-4962-9b52-95a7911ef545 - https://www.gbif.org/occurrence/2643335826; rev. Ó. Sánchez Pedraja Aug 2022); "Herbarium M[artin]. Dürer. / Orobanche arenaria Borkh. / Flora des Grossherz. Hessen / Fundort Sandfelder bei Arheilgen / unweit Darmstadt. Auf Artemisia camp. / Zeit VI. [18]84 / Leg. M. Dürer / Frankfurt a. M. (Deutschland)" (FR 110886, neotype of O. arenaria Borkh. (Uhlich & Gregor 2017)).

Rhineland-Palatinate (DEB): Palatinatus: Dürckheim, Edenkoben ... Prov. Rhenana: Ehrenbreitstein, Hammerstein (Beck 1890: 131, sub O. arenaria).

Sachsen-Anhalt / Saxony-Anhalt (DEE): Roelsdorf prope Halle (Beck 1890: 131, sub O. arenaria).

Thuringia (DEG): "ad Genistae tinctoria radices in colle herbido der Steinklippe. Jun. [near Wendelstein, SE of Rossleben / Roßleben (Until 1815 Roßleben was located in Wendelstein, Saxony)]" (Skočdopolová & Chrtek, 2008: 26) ... "The specimen PR 11757 is selected here as a lectotype. According to the January 2000 revision by German botanist J. Pusch this specimen corresponds to Orobanche arenaria Borkh." (Skočdopolová & Chrtek, 2008: 26)" (Wallroth 1822: 314, loc. class. of Orobanche comosa Wallr.); 1. "O. arenaria Borkh. / Beck [m. Beck]. – 2. "O. comosa mea". – 3. "187 W" (BW 11593030; rev. Ó. Sánchez Pedraja Sep 2016 (Beck 1890: 129 "vidi orig. in herb. Berol."); "Orobanche arenaria, Borkh. / Flora des Kyffhäuserkreis: bei Frankenhausen / Aug [18]91 legi Rich. Schonfelden" (https://data.biodiversitydata.nl/naturalis/specimen/AMD.84029 - https://medialib.naturalis.nl/file/id/AMD.84029/format/large - https://www.gbif.org/occurrence/2513061028; rev. Ó. Sánchez Pedraja Dec 2024); ...auf Artemisia campestris ...  an der Ochsenburg/Kyffhäuser, 9. Juli. 1989 (Uhlich & al. 1995: 40 Abb22!, 41 Abb23!).

Other locations: "Germania: Palatinatus: Dürckheim, Edenkoben; Princ. Badensis: Mannheim, Heidelberg, St. Ilgen; Carlsruhe, Durlach, Wertheim etc.; Darmstadt, Mainz, Heppenheim, Hanau, Prov. Rhenana: Ehrenbreitstein, Hammerstein; Bavaria: München; Prov. Saxonia: Roelsdorf prope Halle; Weimar: Brembach; Silesia: Eulau [Iława Slaska, Warmian-Masurian Voivodeship, Poland]; Brandenburg: Freienwalde, Ruinenburg, Prenzlau, Frankfurt ad fl. Oder, Wriezen; Mecklenburg-Strelitz: Feldberg" (Beck, 1890: 131, sub O. arenaria); "Selten nördliche Oberrheinebene/O-Pfalz; sehr selten SW-Sachsen-Anhalt, N-Thüringen, NO-Brandenburg, Z-Mecklenburg-Vorpommern, Kaiserstuhl, Mainfranken, W-Eifel" (Hassler & Schmitt 2009, photos! [Orobanche in Flora-de: Flora von Deutschland by Thomas Meyer]; accessed Jun 2015).

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Hungary / Hungría / Magyarország (Borbás 1886: 106, loc. class. of O. arenaria f. gigantiflora; Kerner 1874: 46, loc. class. of O. ionantha; Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria)

Pest County / Pest megye: 1. "Orobanche arenaria / Borkhausen / ..., Erd-Ofalu  ... / 29.6.1974". – 2. "Herbarium /Georgii Dominicque SAG / Orobanche arenaria / Borkhausen / sur loess du Sánchegy / Érd-Ófalu près de Budapest / 29.6.1974, leg Gotth 3396" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p00044449 - P 00044449.jpg - https://www.gbif.org/occurrence/437955925 - https://explore.recolnat.org/occurrence/7BABCBDB0EBB4C9A87EED4480109BA4C; rev. Ó. Sánchez Pedraja Jul 2022).

Other locations: "Unter den in Ungarn [Hungary] gesammelten Orobanchen fand sich auch ein Exemplar einer noch nicht beschriebenen Art, welche ich seither auch an mehreren Punkten in den Thälern der Alpen auf Artemisia campestris auffand, und die ich insbesonders in der Nähe von Innsbruck [Tyrol] auf den Angern der sogenannten "Höttinger Aue [Höttinger Au]"..." (Kerner 1874: 46, loc. class. of O. ionantha); "Hungaria: Prope Pressburg [Bratislava, Slovakia], in monte Haglersberg ad lacum Peisonem [Neusiedler See / Fertő tó, Austrian–Hungarian border (next to Jois (Nyulas) in Austria and to Sopron / Ödenburg in Hungary)], prope Adamow com. Neutra [Adamovský les wood near the village of Gbely, Horná Nitra, Slovakia]] (forma 1 cum forma 2 leg. Krzisch), in monte Adlersberg et Blocksberg prope Budam [Budapest] (Ofen), Mézhegy, Grebenac [Grebenac] (Banat) etc." (Beck 1890: 131, sub O. arenaria).

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Iran / Irán / Persia [unconfirmed]

"Bakht.: Kuh Safid, Pau & Vic. [1899, leg. F.M. de la Escalera, det. C. Pau & C. Vicioso]" (Schiman-Czeika 1964: 11, sub O. arenaria).

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Iraq / Irak / Ирак [in error]

Sulaymaniyah Governorate: 1. "Professor C. Haussknecht. It. Orient. 1867 / Phelipaea ramosa Reuter / ad rad. Composit. [?] Pir Omar / Gudrun [Shak-i Pira Magrun / Jabal Birah Magrun / Pira Magroon, Dokan District / Sulaymaniyah District], Kurdist. / dieb.  June // 1870 ... ". – 2. "Orobanche arenaria Borkh. in GBIF" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02982384 - P 02982384.jpg - https://www.gbif.org/occurrence/437106827; rev. Ó. Sánchez Pedraja Sep 2016, sub P. libanotica, syntype of O. coelestis f. confinis (Piwowarczyk & al. 2023: 27, isolectotype)).

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Israel / תישראל / َائِيل‎ / and Palestinian territories [in error]

Judea and Samaria District (incl. West Bank / Yehuda ve-Shomron / Cisjordanie / Cisjordania): Jericho Governorate: 1. "Flora Palaestina / Orobanche ramosa L. / Abundant in deeper soil of wadi dominated / by Malva and Urtica. Malva most frequent host. Auja spring [Al-Auja], north of Jericho. / Semi-steppe and batha vegetation. 35 22' E, 31 58' N / L.J. Musselman 101135 18 February /1987". – 2. "Orobanche arenaria Borkh. / Det.-rev. D. Costa, Junho 1998" (http://data.rbge.org.uk/herb/E00074514 - E00074514.zip - https://www.gbif.org/occurrence/574952690; rev. Ó. Sánchez Pedraja May 2024, sub P. aegyptiaca).

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Italy / Italia / Italien (Reuter 1847: 6, sub Phelipaea arenaria; Beck 1890: 131, sub O. arenaria; Béguinot 1902: 473, sub K. arenaria et sub Kopsia arenaria f. ionantha; Beck 1930: 103, sub O. arenaria; Pignatti 1982: 610, sub O. arenaria; Aeschimann & al. 2004: 284, sub O. arenaria; Pusch & Günther 2009: 31, sub O. arenaria)

Emilia-Romagna [unconfirmed]

Forlì-Cesena (FC): "Phelipanche arenaria (Borkh.) Pomel // Orobanche ramosa L. // Orobancaceae / Forli' (FC), 34 m, mag 2012 (https://www.floraitaliae.actaplantarum.org/viewtopic.php?f=40&t=46931 [215568, 215569, 215577, 215578] by Eugenia Bugni in Acta Plantarum; accessed et rev. Ó. Sánchez Pedraja Jul 2017 [the specimens seem to correspond to P. purpurea, apparently, parasitizing Artemisia vulgaris cf.]).

Reggio Emilia (RE): Canossa (RE), 320 m, giu 2013 (photo 50197_230704 by Pierfranco Arrigoni, sub P. arenaria, in Acta Plantarum; accessed et rev. Ó. Sánchez Pedraja Jul 2017 [the specimens seem to correspond to P. nana aggr. (P. lainzii)).

Lombardia (Pignatti 1982: 610, sub O. arenaria)

Brescia (BS): "BS" (Aeschimann & al. 2004: 284, sub O. arenaria); "Provinz Brescia: in der Ebene südlich Brescia auf Artemisia verlotiorum" (Pusch & Günther 2009: 31, sub O. arenaria [see P. purpurea]); Bompensiero [Villachiara] (BS), mag 2008 ... su Artemisia verlotiorum (https://www.floraitaliae.actaplantarum.org/viewtopic.php?f=40&t=4429 [id=10865 - Phelipanche_arenaria_97a5efd7.jpg, id=10375 - Phelipanche_arenaria_1ed7648f.jpg and id=10377 - Phelipanche_arenaria_361d073b.jpg] by di Franco Giordana, sub P. arenaria, in Acta Plantarum; accessed et rev. Ó. Sánchez Pedraja Jul 2017 [the specimens seem to correspond to P. purpurea [yellowish form]).

Pavia (PV): "Pavese" (Béguinot 1902: 473, sub K. arenaria); "Pavese" (Pignatti 1982: 610, sub O. arenaria).

Sondrio (SO): "SO" (Aeschimann & al. 2004: 284, sub O. arenaria).

Varese (VA): "VA" (Aeschimann & al. 2004: 284, sub O. arenaria).

Piemonte (Reuter 1847: 6, sub Phelipaea arenaria; Beck 1890: 131, sub O. arenaria; Béguinot 1902: 473, sub K. arenaria et Kopsia arenaria f. ionantha; Pignatti 1982: 610, sub O. arenaria)

Cuneo (CN): "Pedemontii subalpinis!" (Reuter 1847: 6, sub Phelipaea arenaria); "Alba ... Alpes Cottiarum [Cottian Alps / Alpes Cottiennes / Alpi Cozie] in vall. Ferora (2 [f. ionantha] leg. Rostan)" (Beck 1890: 131, sub O. arenaria); "nelle A. Marit. [Alpi Marittime]" (Béguinot 1902: 473, sub K. arenaria); "Valle aride alpine dal ... A. Maritt." (Pignatti, 1982: 610, sub O. arenaria); "CN" (Aeschimann & al. 2004: 284, sub O. arenaria); Limone P.te [Limone Piemonte] (CN), 1060 m, lug 2016 (photos 89267 [425272, 425273 and 425270] by Giacomo Bellone in Acta Plantarum; accessed Ó. Sánchez Pedraja Jul 2017).

Torino (TO): "Sabione de Cambiano" (Beck 1890: 131, sub O. arenaria); Torino [45°02'18.1"N 7°07'50.5"E - 45.038369, 7.130699, Roure], Jun 2017 (https://www.inaturalist.org/observations/6509220 - https://www.gbif.org/occurrence/3415581313 by bferrero; accessed  Jul 2022 [on Artemisia sp.]).

Toscana (Pignatti 1982: 610, sub O. arenaria)

Firenze (FI): "presso Firenze lungo l'Arno alle Cascine" (Béguinot 1902: 473, sub K. arenaria); "Firenze alle Cascine" (Pignatti 1982: 610, sub O. arenaria).

Trentino-Alto Adige (Pignatti 1982: 610, sub O. arenaria)

Bolzano – Alto Adige / Bozen – Südtirol (BZ): "Tir. [Tirol]" (Béguinot 1902: 473, sub K. arenaria); "Tir. Magg.-Lugl." (Béguinot 1902: 473, sub Kopsia arenaria f. ionantha); "Valle aride alpine dal Trent." (Pignatti 1982: 610, sub O. arenaria); "BZ" (Aeschimann & al. 2004: 284, sub O. arenaria); "Provinz Bozen: mehrfach im Vinschgau [Val Venosta]" (Pusch & Günther 2009: 31, sub O. arenaria).

Trento (TN): "Orobanche arenaria Borkh. [Phelipanche arenaria (Borkh.) Pomel] - Indicata come probabilmente estinta in Trentino da Prosser (2001), è stata successivamente rinvenuta al Castello di Stenico (M. Merli in Festi & Prosser, 2008) e sulla pendice ca. 1 km a est di Mezzana (2008, ROV, AB & FP)." (Bertolli & Prosser 2011: 275, sub O. arenaria); "Provinz Trento: am Schloss Sténico westlich Trento ... Trient [Trento], Revò in Val di Non [Nonsberg] und Riva..." (Pusch & Günther, 2009: 31, sub O. arenaria); Phelipanche arenaria (Borkh.) Pomel / Trentino (TN), 670 m, giu 2007 ... Specie rarissima parassita su Artemisia campestris ... specie molto molto rara e verosimilmente per il Trentino è la sola stazione vitale (photos 267408 et 267409 [58391_267408] by Marco Merli in Acta Plantarum; accessed et rev. Ó. Sánchez Pedraja Jul 2017).

Valle d'Aosta / Vallée d'Aoste (AO): Farys-Chambave [Valle d'Aosta] (AO), 540 m, mag 2016 (photos http://www.actaplantarum.org/galleria_flora/galleria1.php?id=2104 [87120_414088, 87120_414116] by Mauro Ottone in Acta Plantarum; accessed Ó. Sánchez Pedraja Jul 2017); Quart (AO), 650 m, mag 2012 (photos 38255_177881, 38255_177878 et 38255_177880 by Claudia Ganz in Acta Plantarum; accessed Ó. Sánchez Pedraja Jul 2017); "Phelipanche arenaria (Borkh.) Pomel. Regione Valle d'Aosta, 600 m s.l.m., Maggio 2016" (photos 9518-0-27638200-1464511229, 9518-0-46967800-1464511261 et 9518-0-80711200-1464511291 by Giomalvi in Funghi italiani 92924 - A.M.I.N.T.; accessed Ó. Sánchez Pedraja Jul 2017); "Scoperta di recente a Gressan sulla Côte de Gargantua, 740 m (Bovio & Broglio 2013 - AO, FI, Domina det.) [AO-N.SFV-2905: Gressan, 2013 - Domina det.]" (photos 5430.0230-P.jpg, 5430.0230-01.jpg and 5430.0230-03.jpg by M. Broglio, sub Phelipanche schultzi1 1645-102 in 2018 Société de la Flore Valdôtaine; accessed et rev. Ó. Sánchez Pedraja Sep 2018 [the photos correspond to P. arenaria]); Aosta (photos 5430.0030-03.jpg, 5430.0030-06.jpg and 5430.0030-05.jpg by M. Broglio, sub Phelipanche purpurea 1306-102 in 2018 Société de la Flore Valdôtaine; accessed et rev. Ó. Sánchez Pedraja Sep 2018 [the photos correspond to P. arenaria]); "Collina di Chambave. Lungo la strada per Chandianaz, altitudine 530 m, 20 maggio 2016, M. Bovio, M. Broglio, F. Secchi, L. Secchi et al. - G. Domina conferma det. (obs., fot.)." (Bovio 2016: 138, sub O. arenaria); "Segnalata nel cuore della media valle, nella bassa e media V. di Cogne ! e in bassa Valgrisenche a Lolair (Bovio, Broglio & Trompetto); indicata da circa 530 m, sopra Chambave (Bovio, Broglio, Secchi F., Secchi L. et al., 2016 - fot. !; Domina conf. det.) fino a 1400 m, presso Cogne (Bovio, Soldano & Trompetto 2011 - AO, Domina det.).", AO-N.SFV-2727: V. Cogne, 2011 - Domina det. (photos 5430.0020-P.jpg, 5430.0020-01.jpg and 5430.0020-03.jpg by M. Broglio, sub P. arenaria 1305-102 in 2018 Société de la Flore Valdôtaine; accessed et rev. Ó. Sánchez Pedraja Sep 2018 !).

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Kazakhstan / Қазақстан / Qazaqstan / Казахстан / Kazajistán

Aktobe Region Ақтөбе облысы / Aqtöbe oblysy / Актюбинская область: "Bayganinskiy [Bayganin District / Байғанин ауданы, 46°24'48.0"N 57°06'30.5"E 46.413333, 57.108472], 24 May 2024" (https://www.inaturalist.org/observations/218361137 by https://www.inaturalist.org/people/wolfgang_rv, sub P. arenaria, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja May 2024, sub O. amoena, apparently on Artemisia).

Atyrau Region / Атырауская область / Атырау облысы / Гурьевская область: "Phelipaea coerulea / Inder [Ozero Inder / Inder Koli / Inder See / Inderskoye Ozero / Lake Inder / Индерское озеро] Nº 3 Claus / Herb. Al. de Bunge" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02982379 - P 02982379.jpg - hb. Cosson - http://www.gbif.org/occurrence/438113031 - https://explore.recolnat.org/occurrence/1261F22E866541FF9FDE5F2A8B3A1880; rev. Ó. Sánchez Pedraja Sep 2015 [the specimen seems to correspond to P. arenaria]); Западный Тянь-Шань и Каратау (Казахстан) [Western Tien Shan and Karatau (Kazakhstan)] (https://plant.depo.msu.ru/open/public/item/MW0878088 - MW 0878088; rev. Ó. Sánchez Pedraja May 2018 [? see P. pallens]); "Soviet Central Asia: Aralo-Caspian [Aral-Caspian Depression / Арало-Каспийская низменность] (northwestern part)" (Novopokrovskii & Tzvelev, 1958: 53[49], sub O. arenaria).

West Kazakhstan Region / Батыс Қазақстан облысы / Западно-Казахстанская область: 1. "Phelipaea longiflora C.A.M. / ad oppid. Iuder [Iuder, Aider. A mountain and a Salt lake of the same name in the neighbourhood of the river Ural. The lake belongs to the Cossack settlements of the Ural (Oral / Орал / Uralsk / Уральск)]". – 2. "Имперaторckаго Моcковскаго Университета [Imperial Moscow University] / Gremiatschensky, S.J. Die Pflanzen der Bukejevvschen / Kirgisen-Horde. 1855. / (Ryn-Peski Sanddünen [Ryn Desert / Рын-пески / Нарын-Құм / Naryn-Qumt], Gross. Bogdo, Bisch-tau, Jakschibisch-tau Arsagar, Baskuntschatskoje-See [Astrakhan steppe / Kalmyk Steppe] et сеt.)". – 3. "Orobanche arenaria Borkh. / 29.VIII.1949 / Det. I. Novopokrovsky" (https://plant.depo.msu.ru/open/public/item/MW0514043 - https://plant.depo.msu.ru/open/public/item/MW0514043/img/0.jpg?original=2904dadb-71b9-441d-a832-aa8098839291 - MW 0514043.jpg - https://www.gbif.org/occurrence/1697663136; rev. Ó. Sánchez Pedraja May 2024, sub P. arenaria); 1. "Orobanche arenaria MB. / mont Juder [Iuder, Aider. A mountain and a Salt lake of the same name in the neighbourhood of the river Ural. The lake belongs to the Cossack settlements of the Ural (Oral / Орал / Uralsk / Уральск)]". – 2. "Имперaторckаго Моcковскаго Университета [Imperial Moscow University] / Gremiatschensky, S.J. Die Pflanzen der Bukejevvschen / Kirgisen-Horde. 1855. / (Ryn-Peski Sanddünen [Ryn Desert / Рын-пески / Нарын-Құм / Naryn-Qumt], Gross. Bogdo, Bisch-tau, Jakschibisch-tau Arsagar, Baskuntschatskoje-See [Astrakhan steppe / Kalmyk Steppe] et сеt.)" (https://plant.depo.msu.ru/open/public/item/MW0514039 - MW 0514039.jpg - https://plant.depo.msu.ru/open/public/item/MW0514039/img/0.jpg?original=f5f0796d-fdcb-49c2-93af-bfa6d425145f - https://www.gbif.org/occurrence/1697663129; rev. Ó. Sánchez Pedraja May 2024, sub O. cumana).

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Moldavia / Moldova / Moldau / Молдавия (Gejdeman 1986)

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Morocco / المغرب‎, / Marruecos / Marokko (Jahandiez & Maire 1934: 694, sub O. arenaria)

Drâa-Tafilalet

Ouarzazate Province: "Siroua [Adrar Sirwa / Adrar Siroua / Jbel Sirwa (Anti-Atlas)], 12 Jun 2025" (https://www.inaturalist.org/observations/290770331 by https://www.inaturalist.org/people/dincastro [Dinis Castro], sub P. arenaria, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Nov 2025 [!]).

Fès-Meknès

Ifrane Province: 1. "Plantae Maroccanae Exsiccatae / a Herbario Vadensi distributae / J.J.F.E. et P.A.W.J. de Wilde et J. Dorgelo collectores / Orobanche sp. / Loc.: Dayet Aouaoua. 17 km from / Ifrane. [33°39'11"N, 5°2'30"W (= 33.653333, -5.041667), Middle Atlas] / No 2611 Die May 28, 61 [1961] / ...". – 2. "Orobanche arenaria Borkh. ? / Det. F.M. Muller, 1982 Herbarium Vadense (WAG)" (https://data.biodiversitydata.nl/naturalis/specimen/WAG.1141172 - WAG 1141172.jpg - https://www.gbif.org/occurrence/2517457982; rev. Ó. Sánchez Pedraja Nov 2018 [!]).

Marrakesh-Safi Region (Marrakech-Tensift-El Haouz) [unconfirmed]

Al Haouz Province: "Oukaimden [31°12'16.7"N 7°51'08.1"W 31.204650, -7.852260], 17 Jun 2025" (https://www.inaturalist.org/observations/293378608 by https://www.inaturalist.org/people/dincastro [Dinis Castro], sub P. arenaria, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Nov 2025, sub P. reuteriana [hairy plant]).

Oriental

Oujda-Angad- Berkane- Taourirt Provinces: "Sur les Artemisia ... R. Beni Snassen [Béni-Snassen / Aït Iznassen / Monts des Beni Snassene, N 34° 48' 0'' W 2° 20' 0''] (Boigey 93)" (Jahandiez & Maire 1934: 694, sub O. arenaria).

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North Macedonia / Северна Македонија / Macedonia del Norte / Macédoine du Nord / Nordmazedonien / Северная Македония (Teofilovski & Mandzukovski 2023: 97-98, sub P. arenaria; Nikolov 2024: 27, sub P. arenaria)

Eastern Statistical Region / Источен Регион

Štip Municipality / Општина Штип: "Štip [Shtip / Штип], above Tri Češmi [Три Чешми] village, open dry place, 515 m, 41.773109°N, 22.116803°E, 13.6.2021 [41°46'23.2"N 22°07'00.5"E 41.773109, 22.116803], leg. A.T. & D.M. [Aco Teofilovski & Dejan Mandzukovski], det. A.T. ... most probably parasitize on Achillea coarctata Poir." (Teofilovski & Mandzukovski 2023: 97-98, sub P. arenaria); "Štip [Shtip / Штип], v. Tri Češmi [Три Чешми], 02.06.2022" (photographies by Z. Nikolov [Z. Nikolov, pers. comm., May 21, 2024]).

Skopje Statistical Region / Скопски Регион

Čučer-Sandevo Municipality / Општина Чучер-Сандево: "Skopska Crna Gora Mt. [Skopska Crna Gora / Karadak Mountains / Скопска Црна Гора] (Crn Kamen) (Grupče 1958, sub Orobanche arenaria)" (Teofilovski & Mandzukovski 2023: 97, sub P. arenaria).

Skopje / Скопје: "Skopje (Osoj [Осој]) (Matveeva 1968, sub O. arenaria)" (Teofilovski & Mandzukovski 2023: 97, sub P. arenaria).

Note: "Phelipanche arenaria (Borkh.) Pomel (Grupče, 1958; Matvejeva, 1968; Teofilovski & Mandzukovski, 2023)" (Nikolov 2024: 27).

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Poland / Polska / Polonia / Polen (Beck 1890: 131, sub O. arenaria; Piwowarczyk & al. 2010: 44)

Silesia: Eulau [Iława Slaska, Warmian-Masurian Voivodeship] (Beck 1890: 131, sub O. arenaria); "The species is very rare in Poland, it has so far been confirmed or reported as occurring at only four localities: in Lower Silesia (Szczęśniak 2003) and in the Niecka Nidziańska basin (Piwowarczyk and Przemyski 2010). It was recorded at one locality in the Middle Vistula Gap in the study area (Fig. 2). It flowers at the end of June and the beginning of July. It parasitizes Artemisia campestris. List of localities: MPW: Męćmierz – the N and NEfacing slope of a small dry valley, ca. 1.5 km SW of the village, a fallow field with dominant vegetation of the class Artemisietea vulgaris (>100), FE2323, 51°18’07”N, 21°53’54”E, vid. P. Chmielewski, 22.04.2009, leg. R. Piwowarczyk, 26.07.2009 (KTC)." (Piwowarczyk & al. 2011: 40).

"25.06.2007, Ponidzie; Renata Piwowarczyk" (photos in Atlas roślin; accessed Dec 2014).

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Portugal (Guimarães 1904: 82, sub O. arenaria; Beck 1930: 103, sub O. arenaria; Aedo & al. 1997: 338, sub O. arenaria)

North Region / Região Norte

Minho [historical province of Portugal] (Mi): Viana do Castelo District, Caminha, "Praia do Moledo [Moledo do Minho] (sobre a Arth. crith. L. [Artemisia crithmifolia L.], R. da Cunha!)" (Guimarães 1904: 82, sub O. arenaria; Sampaio 1947: 499 sub Kopsia purpurea; Aedo & al. 1997: 338, sub O. arenaria!); "Viana do Castelo: Cabedelo, nos areais sobre Artemisia crithmifolia, 29TNG11, 24-VI-1959, A. Rozeira, K. Koepp & G. Costa, PO 21359" (Domingues de Almeida 2003: 59, sub O. purpurea Jacq.); "Viana do Castelo: dunas de Amorosa, nos areais, 29TNG11, 8-VI-1961, A. Rozeira, K. Koepp & G. Costa, PO 21360" (Domingues de Almeida, 2003: 59, sub O. purpurea Jacq.); Braga District, Esposende Municipality, "26-6-1972 / Ofir: no pinheiral à beira / mar / col. Rozeira, Araújo e Serra" (MA 499870, sub O. purpurea; rev. ut possibly O. purpurea ?, M.J.Y. Foley 1998; rev. ut O. arenaria, A. Pujadas II-2002; rev. Ó. Sánchez Pedraja 8 Nov. 2007); ib., PO 21361 (Domingues de Almeida, 2003: 59, sub O. purpurea Jacq.); concelho de Esposende, "Erva-toira-das-areias (Orobanche arenaria) [photo!]" (J. Silva, 28 de Maio de 2008, Flora Dunar do Litoral Norte).

Douro Litoral (DL): "Póvoa de Varzim: Aguçadora, in arenosis maritimis hosp. Artemisia crithmifolia, 10 m s.m., 29TNF18, 10 m., 1-VII-1960, A.R. Pinto da Silva, B.V. Rainha & M. da Silva 6644" (LISE 88538) (Domingues de Almeida, 2003: 59, sub O. arenaria); "Vila do Conde: Mindelo, areais, 19-VI-1984, 29TNF27, Barreto & A. Serra PO 53030" (Domingues de Almeida 2003: 59, sub O. purpurea Jacq.).

Centro Region / Região Centro

Beira Litoral [historical province of Portugal (BL): Coimbra District, "Figueira da Foz: dunas, Leirosa (F. Loureiro!)" (Guimarães, 1904: 82, sub O. arenaria); Coimbra District, Figueira da Foz, "Quiaios [Praia de Quiaios] (na Arth. crith. L. [Artemisia crithmifolia L.], A. de Carvalho! que foi quem, em 31 de julho de 1866, pela primeira vez, em Portugal, colheu esta especie)" (Guimarães 1904: 82, sub O. arenaria).

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Romanie / România / Rumanía / Rumänien / Румыния (Beck 1890: 131, sub O. arenaria; Beck 1930: 103, sub O. arenaria)

Cluj County: "Szenafü bei Klausenburg [Cluj-Napoca]" (Beck 1930: 103, sub O. arenaria); "Romania, județul Cluj. Cluj Napoca, Fânaţele Clujului, dry steppes on south-facing slopes; 450-480 m s. m., coordinates: 46°49'52"N & 23°38'13"E ; Alt. 450 m - 480 m, 2017-06-09, Chytrý, M. et al. Iter romanicum anno 2017 108" (BRNU 655570 - https://brnu.jacq.org/BRNU655570 - https://www.jacq.org/detail.php?ID=1199939  - https://www.gbif.org/occurrence/2382580105 [n.v.]).

Sibiu County: 1. "Orobanche arenaria Borkh. / Trionychon arenarium Sckur / Auf Sandhügeln bei Hammersdorf [Gușterița] / auf Artemisia campestris L. / 8 July 1891 / det. Dr. Beck". – 2. "Orobanche aegyptiaca Pers. / forma ... serotina Hausk." (E 01005562 - hb. Dörfler - https://data.rbge.org.uk/herb/E01005562 - https://www.gbif.org/occurrence/3312568578; rev. Ó. Sánchez Pedraja Jul 2022); "Hammersdorf [Gușterița]" (Beck 1890: 131, sub O. arenaria), " Hammersdorf [Gușterița]" (Beck 1930: 103, sub O. arenaria).

Tulcea County / Județul Tulcea: "Bulgaria: prope Tulscha [Tulcea] Dobrutschae [Dobruja] leg. Sintenis" (Beck 1890: 131, sub O. arenaria).

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Russia / Россия / Rusia / Russland / Russie (Koch 1849: 661; Beck 1890: 130, loc. class. of O. arenaria f. sareptana; Beck 1930: 103, sub O. arenaria; Novopokrovsky & Tzvelev 1958: 53[49], sub O. arenaria, sub O. arenaria; Tzvelev 1981 [2001]: 447; sub P. laevis in Plantarium; accessed Feb 2012 et Jun 2017; Uhlich & Rätzel 2024: 6, sub P. arenaria var. sareptana)

Central Federal District / Центральный федеральный округ

Belgorod Oblast / Белгородская область: 1. "... / Orobanche purpurea Jacq. / ... Volgograd Prov., Frolovo dist. [Frolovsky District] ... Archeda ... / on sand / 49°46'00.0"N 43°22'00.0"E - 49.766667, 43.366667 / 25.VII.1993 Coll.: I. Shantser ...". – 2. "Orobanche laevis L. ... 16.03.2021" (MHA 0162489 - https://plant.depo.msu.ru/open/public/item/MHA0162489 - https://plant.depo.msu.ru/open/public/item/MHA0162489/img/0.jpg?gbif - https://www.gbif.org/occurrence/3345589158; rev. Ó. Sánchez Pedraja Aug 2022); Новооскольский район [Novooskolsky District] 9.07.2000 А.Н. Федотов [A.N. Fedotov]" (https://plant.depo.msu.ru/open/public/item/MW0514028 - MW 0514028.jpg; rev. Ó. Sánchez Pedraja May 2018); "... / Orobanche laevis L. / Старооскольский район [Starooskolsky District]... / c. Дмитриевка [Dmitrievka] / 51.246805, 38.181965 - 51°14'48.5"N 38°10'55.1"E / ... / 04.VII.2017 / Н.М. Решетникова [N.M. Reshetnikova] et al." (MHA 0008828 - https://plant.depo.msu.ru/open/public/item/MHA0008828 - https://plant.depo.msu.ru/open/public/item/MHA0008828/img/0.jpg?gbif - https://www.gbif.org/occurrence/3345591747; rev. Ó. Sánchez Pedraja Aug 2022).

Kursk Oblast / Курская область: "Цветущее растение. Курская обл., Горшеченский р-н [Gorshechensky District], окр. дер. Кунье, степь на склоне балки. 06.07.2014" (http://www.plantarium.ru/page/image/id/449458.html and plantarium.ru/page/image/id/449460 by Евгений Скляр [Evgenij Sklyar] 2016 in Plantarium; accessed June 2017).

Lipetsk Oblast / Липецкая область: Volovsky District / Воловский район, 10.07.1982 (https://plant.depo.msu.ru/open/public/item/MW0514029 - MW 0514029.jpg; rev. Ó. Sánchez Pedraja May 2018).

North Caucasian Federal District / Северо-Кавказский федеральный округ

Karachay-Cherkessia / Карачаево-Черкесская Республика / Къарачай-Черкес Республика: "Herbarium Universitatis Mosquensis (MW) / Orobanche caesia Reichb. / Карачаево-Черкесская, / ... Nº 7335 15 июня 2010 г." (https://plant.depo.msu.ru/open/public/item/MW0723412 - MW 0723412; rev. Ó. Sánchez Pedraja May 2018).

Dagestan (Republic of) / Республика Дагестан: "Im südlichen Daghestan, c. 800' hoch" (Koch 1849: 661, sub P. arenaria Borkh.); "in Dagestan: bei Petrowsk [Петровск, Makhachkala / Махачкала] an sandigen Stellen an der Küste des Kaspischen Meeres (Fedtschenko, Lipsky!)" (Beck 1930: 104, sub O. arenaria); Herbarium Universitatis Mosquensis (MW) O. arenaria Borkh. (https://plant.depo.msu.ru/open/public/item/MW0723408 - MW 0723408.jpg; rev. Ó. Sánchez Pedraja May 2018); "Orobanche arenaria Borkh. / Darewan Дербентский район [Derbentsky District], ces Геджух [Gedzhukh] / ... /  1970  A. Gadzhiomarov" (https://plant.depo.msu.ru/open/public/item/MW0723409 - MW 0723409.jpg - https://www.gbif.org/occurrence/1697840861; rev. Ó. Sánchez Pedraja May 2024, sub P. purpurea); "Цветущее растение. Дагестан, Ахтынский р-н [Akhtynsky District], окр. с. Ахты. 30.06.2015." (http://www.plantarium.ru/page/image/id/453501.html by Юсиф Саркаров [Yousif Sarkarov] 2016; растение определил(а) Андрей Ковальчук in Plantarium; accessed Jun 2017 [rev. ut Phelipanche caesia (Rchb.) Soják, inflorescence arachnoid-villous!]); "Цветущие растения (паразитируют на Artemisia sp.). Дагестан, Дербентский р-н [Derbentsky District], побережье Каспийского моря в 4 км к северо-востоку от пгт Мамадкала [Mamedkala], частично закрепленные пески. 14 июня 2018 г. (https://www.plantarium.ru/page/image/id/575758.html by Александр Фатерыга [Alexander Fateryga] 2018 in Plantarium; accessed Jan 2022); Респ. Дагестан [43°00'05.8"N 47°13'37.0"E - 43.001601, 47.226947, near Sarykum / Sary-Kum / Сары-Кум (Kumtorkalinsky District / Кумторкалинский район)], 1 Jun 2021 (https://www.inaturalist.org/observations/81529697 by Anna Nikolaeva in iNaturalist; accessed Jul 2022).

Kabardino-Balkar Republic / Кабардино-Балкарская Республика: "Кабардино-Балкария, Верхняя Балкария [Kabardino-Balkaria, Upper Balkaria]" (http://www.plantarium.ru/page/image/id/114791.html by Сергей Банкетов, 2011; растение определил(а) Евгений Киреев, sub P. laevis in Plantarium; accessed Feb 2012).

North Ossetia-Alania Republic / Республика Северная Осетия-Алания (former North Ossetian Autonomous SSR / Северо-Осетинская АССР): 1. Herbarium Universitatis Mosquensis (MW) / Северо-Осетинская АССР, бассейн р. Ардон, / южный склон Скалистого хребта над пос. / Мизур (левый борт Алагирского ущелья) [Nort Ossetian ASSR, the Ardon Riber basin ... above village of Mizur ... 19 7 A.M.Амирханов [A.M. Amirkhanov] ". – 2. "Orobanche pulchella (C.A. Mey.) Novopokr. [= O. caucasica G. Beck ] / 233 / 1977 Teste Амирханов [Amirkhanov]". – 3. "Herbarium Universitatis Mosquensis (MW) / Orobanche pulchella / (C.A. Mey.) Novopokr. / [= O. caucasica G. Beck ] / 233 ... Сухие щебн. склоны ... 1000-1500 м ... /  ... 29.VII.76[1976] ..." (https://plant.depo.msu.ru/open/public/item/MW0717040 - MW 0717040 - https://www.gbif.org/occurrence/1697834525; rev. Ó. Sánchez Pedraja Nov 2017 [the three specimens seem to correspond to P. arenaria]).