Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors

Accepted Name:

Orobanche pubescens d’Urv., Enum. Pl. Ponti-Eux.: 76 (1822 [Apr. 1822]) [et Mém. Soc. Linn. Paris 1: 332 (1822)] (cf. Reuter, 1847: 27; Reichenbach fil., 1862: 101; Beck, 1902: 454; Beck, 1930: 183; Gilli, 1982: 15; Domina & Raab-Straube, 2010)

Synonyms:

≡ Orobanche minor var. pubescens (d'Urv.) R.D. Meikle, Fl. Cyprus 2: 1242 (1985) (cf. Boulos, 2002: 94)

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= Orobanche grandisepala F.W. Schultz, Flora (Regensb.) 26: 128 (1843). Ind. loc.: "Ad Naupliam leg. Cl. Fraas" [Nafplion / Nauplia / Navplion and Corinth / Korinth; Greece]

= Orobanche tridentata F.W. Schultz, Flora (Regensb.) 26: 128 (1843) ["O. tridentata F.G. Schultz 1834 in litt."], nom. illeg. Syn.: = Orobanche grandisepala F.W. Schultz, Flora (Regensb.) 26: 128 (1843)

= Orobanche arcuata F.W. Schultz, Flora (Regensb.) 26: 129 (1843). Ind. loc.: "In Attica Cl. Fraas" [Attica, Greece]. Obs. "pilis albidis sparsim lanuginosa" (cf. Beck, 1890: 240; Beck, 1922: 455)

= Orobanche versicolor F.W. Schultz, Flora (Regensb.) 26: 129 (1843). Syn: ≡ Orobanche versicolor f. typica Beck in Biblioth. Bot. 19: 239, t. 4 f. 75 (1890), nom. illeg. Ind. loc.: "In Crysanthemo coronopifolio ad Naupliam legit Clarissim. Spruner et prope Corynthum Cl. Fraas" [Nafplion / Nauplia / Navplion and Corinth / Korinth; Greece] (cf. Jordan, 1846: 226 "O. pubescens d’Urv.—versicolor Schultz ... Marseille ... sur le Crepis bulbosa Cass."; Reuter, 1847: 27; Reichenbach fil., 1862: 101; Gilli, 1982: 15 "Nauplia, Spruner (holo. P).")

= Orobanche chrysanthemi F.W. Schultz, Flora (Regensb.) 26: 129 (1843) ["O. Chrysanthemi F.G. Schultz 1834 in herb."], nom. illeg. Syn.: = Orobanche versicolor F.W. Schultz, Flora (Regensb.) 26: 129 (1843)

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= Orobanche villosiflora F.W. Schultz, Flora (Regensb.) 28: 737, 740 (1845), nom. illeg. [pro syn.] (cf. F.W. Schultz, 1845: 737, 740 "O. villiflora ... O. villosiflora ... Meiner Diagnose ist aber "floribus vilosis"; Beck, 1930: 183; (IK) view IPNI entry)

= Orobanche villiflora K. Koch ex F.W. Schultz, Flora (Regensb.) 28: 737, 740 (1845), nom. illeg. [ pro syn.]. Syn.: = Orobanche villosiflora F.W. Schultz, Flora (Regensb.) 28: 737, 740 (1845)

= Orobanche vitalbae Bertol., Fl. Ital. 6: 441 (1846) ["Vitalbae"]. Ind. loc.: "Perenn. Habui ex Etruria in herbidis di Campiglia a Prof. Petro Savio, qui reperit parasiticam supra Clematidem Vitalbam L. Floret Junio [Campiglia Marittima, Toscana / Tuscany, Italy / Italia]" (cf. Bertoloni, 1846: 441 "Corolla ... tubulosa, alba, extus hirsuta..."; Beck, 1890: 239; Beck, 1930: 184, 186; Uhlich & al., 1995: 125)

= Orobanche fragrans Griseb., Spicil. Fl. Rumel. 2 : 58 (1844) [nom. illeg.], non Orobanche fragrans Raf. in Amer. Monthly Mag. & Crit. Rev. 1(6): 440 (1817 [Oct 1817]) nec Orobanche fragrans W.D.J. Koch in Röhl., Deutschl. Fl., ed. 3, 4: 463 (1833) (cf. Reuter, 1847: 27)

= Orobanche arachnoidea F.W. Schultz, Linnaea 19: 26 (1846) ["msc. No 1"], nom. nud. (cf. K. Koch, 1849: 669 "ohne Diagnose"; Beck, 1890: 238)

= Orobanche chrysanthemi F.W. Schultz ex Reut. in DC., Prodr. 11: 27 (1847), pro syn. (cf. F.W. Schultz, 1843: 26; Beck, 1890: 239; Beck, 1930: 186)

= Orobanche villiflora K. Koch ex Reut. in DC., Prodr. 11: 27 (1847) ["mss."], pro syn. (cf. Beck, 1890: 238; Beck, 1930: 183 "(vidi spec. authent.) in litt. et herb.")

= Orobanche villosa F.W. Schultz ex Reut. in DC., Prodr. 11: 27 (1847) ["in litt. 1831"], pro syn. (cf. Reichenbach fil., 1862: 101; Beck, 1890: 238; Beck, 1902: 454; Beck, 1930: 183)

= Orobanche superficialis Griseb. ex Reut. in DC., Prodr. 11: 28 (1847) ["Griseb.! mss. in h. Boissier"], pro syn. (cf. Beck, 1930: 185)

= Orobanche arachnoidea F.W. Schultz ex K. Koch, Linnaea 22: 669 (1849). Syn.: ≡ Orobanche arachnoidea F.W. Schultz, Linnaea 19: 26 (1846) ["msc. No 1"], nom. nud. Ind. loc.: "Von Dr. Thirke aus der Umgegend von Brussa und von Rögner aus der Krim erhalten" [Bursa, Marmara Region, Turkey and Krim / Crimea, Ukraine (cf. Reuter, 1847: 27 "Schultz ex C. Koch [K. Koch] in Thirke pl. or. exsicc.!"; K. Koch, 1849: 669; Beck, 1930: 184 "in Linnaea XIX. 26 (ex diagn. et XXII. (1849) 669; vidi orig. e manu C Kock! [K. Koch]"; Gilli, 1982: 15; (IK) view IPNI entry)

= Orobanche squalida Steven in Bull. Soc. Nat. Mosc. 30: 354 (1857) (cf. Novopokrovskii & Tzvelev, 1958[2000]: 125)

= Orobanche livida Halácsy in Österr. Bot. Z. 47: 97 (1897), non Orobanche livida Sendtn. ex Rchb. fil., Icon. Fl. Germ. Helv. 20: 104 (1862 [22 Dec 1862]) (cf. Beck, 1930: 183 "sec. Halácsy")

= Orobanche campuliflora (Lojac.) Lojac., Fl. Sic. 2(2): 170 (1904) (f. Beck, 1890: 239; Uhlich & al., 1995: 125; Domina & Arrigoni, 2007: 127)

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= Orobanche minor var. adenostyla Halácsy in Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 61: 28 (1894), non Orobanche minor var. adenostyla Vis., Fl. Dalmat. 2: 179 (1847 [10-13 Nov. 1847]) (cf. Beck, 1902: 455; Beck, 1930: 186)

= O. pubescens var. campuliflora Lojac. in Naturalista Sicil. 2(4): 82 (1883). Syn.: ≡ O. campuliflora (Lojac.) Lojac., Fl. Sic. 2(2): 170 (1904); ≡ Orobanche versicolor f. campuliflora (Lojac.) Beck in Biblioth. Bot. 19: 239 (1890) (cf. Beck, 1930: 186; Domina & Mazzola, 2007: 225; Domina & Arrigoni, 2007: 127). Ind. loc.: "In sylvaticis fruticetis maritimis, specimen unicum inveni in Palermo M. Gallo die 10 majo 1881". Ic.: Naturalista Sicil. 1: t. 9, f. 1 (cf. Domina & Mazzola, 2007: 225). Obs. Holotype: PAL 43240 (cf. Domina & Mazzola, 2007: 225)

= O. pubescens var. pallida Reut. in DC., Prodr. 11: 27 (1847) ["ß pallida"]. Syn.: ≡ Orobanche superficialis Griseb. ex Reut. in DC., Prodr. 11: 28 (1847), pro syn.; = Orobanche versicolor f. homochroa Beck in Biblioth. Bot. 19: 238 (1890); = Orobanche versicolor f. monochroa Beck in Halácsy, Consp. Fl. Graec. 2: 455 (1902). Ind. loc.: "In Macedoniâ (Griseb.!)"

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= Orobanche versicolor f. campuliflora (Lojac.) Beck in Biblioth. Bot. 19: 239 (1890)

= Orobanche versicolor f. corcyrensis Beck in Repert. Spec. Nov. Regni Veg. 18: 40 (1922). Ind. loc.: "Corcyra ad castellum, m. Aprili (Bornmüller)" [Nísos Kérkira / Corfu / Corcyra, Greece]

= Orobanche versicolor f. homochroa Beck in Biblioth. Bot. 19: 238 (1890)

= Orobanche versicolor f. lanifera Beck in Biblioth. Bot. 19: 239 (1890). Ind. loc.: "Cum typo hinc inde"

= Orobanche pubescens f. luteostigma Gilli in Feddes Repert. 82(6): 399 (1971). Ind. loc.: Turkey (view IPNI entry)

= Orobanche versicolor f. macrophyllon Beck in Biblioth. Bot. 19: 239 (1890). Syn.: = Orobanche minor var. adenostyla Halácsy in Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 61: 28 (1894), non Orobanche minor var. adenostyla Vis., Fl. Dalmat. 2: 179 (1847 [10-13 Nov. 1847]) (cf. Beck, 1902: 455)

= Orobanche versicolor f. monochroa Beck in Halácsy, Consp. Fl. Graec. 2: 455 (1902)

= Orobanche versicolor f. pseudobarbata Beck in Biblioth. Bot. 19: 239 (1890). Ind. loc.: "Rarius cum typo"

= Orobanche versicolor f. redundans Beck in apud Bornmüller in Mitt. bot. Ver. Thüring. 24: 96 (1908). Ind. loc.: "Lydien: auf dem Berge Sipylos ober Magnesia, bei 200—300 m Seehöhe, im Mai 1906 (Bornmüller!)."

= Orobanche versicolor f. typica Beck in Biblioth. Bot. 19: 239, t. 4 f. 75 (1890), nom. illeg. Syn: ≡ Orobanche versicolor F.W. Schultz, Flora (Regensb.) 26: 129 (1843)

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Ind. loc.: "Vulgaris in collibus siccis insulae Meli. Aprili floret" [Milos / Melos (cf. Reuter, 1847: 27; Beck, 1930: 184), Cyclades islands, Greece].

Typus / Type:

Designated by:

Host: Usually on Compositae and Umbelliferae species (e.g., Sonchus bulbosus (L.) N. Kilian & Greuter [≡ Aetheorhiza bulbosa (L.) Cass.], Glebionis segetum (L.) Fourr. [≡ Chrysanthemum segetum L.]) and perhaps on others families too (cf. Annotated Checklist of Host Plants of Orobanchaceae, accessed 24 August 2009).

Comments: "Since Orobanche pubescens Gilib. (1781, 1782) [view IPNI entry] is invalid (See Appendix V: 334. ICBN, 1994 [Vienna Code, Appendix VI Opera utique oppressa]), O. pubescens d’Urv. (1822) is not a later homonym" (cf. Novopokrovskii & Tzvelev, 1958[2000]: 125).

Documentos Jard. Bot. Atlántico (Gijón) 3: 61 f. 4 ["Phylogenetic relationship"], 65 (2005).

Documentos Jard. Bot. Atlántico (Gijón) 6: 13 f. 2 ["Phylogenetic relationships"].

Chromosomatic number:

Ic.: a, Reichenbach fil. (1862: t 180); b, Beck, 1890:  t. 4 f. 75 (sub O. versicolor); c, habit (on Glebionis segetum; Port-Vendres, Cap Béar, France [by Virgile Noble]); d, flowers (ib.); e, habit (on Aetheorhiza bulbosa; Marseille sur l’île de Maïre, France [by Virgile Noble]);d, whitish flowers (Samariá Gorge, Crete, Greece [by Henri Michaud]); f, flowers white tinged with purple (ib.).

Distribution: distribution (sec. Domina & Raab-Straube, 2010).

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Specimens examined and literature records.

Albania: (cf. Zerny in Hayek, 1924: 166; Barina & Pifkó, 2008: 460).

Bulgaria: (cf. Grisebach, 1844: 58; Beck, 1930: 184; Stoyanov, 2009: 355).

Crimea: (cf. Novopokrovskii & Tzvelev, 1958[2000]: 126).

Croatia / Croacia: (cf. Beck, 1930: 184; Greuter & al., 1989: 261).

Cyprus / Chipre: (cf. Beck, 1890: 240; Meikle, 1985: 1242).

Egypt / Egipto: (cf. Beck, 1930: 185; Boulos, 2002: 94; Greuter & al., 1989: 261).

France / Francia: (cf. Jordan, 1846: 226; Reynier, 1913; Crouzet & al., 2009: 60).

Corsica / Córcega: (cf. Habashi & Jeanmonod, 2008: 60).

Georgia: (cf. Beck, 1890: 240).

Greece / Grecia: (cf. Dumont d'Urville, 1822: 76; F.W. Schultz, 1843: 128, 129; Reuter, 1847: 27; Beck, 1930: 184).

Crete / Creta: (cf. Beck, 1890: 240; Greuter & al., 1989: 261; P.C. Brouwer, Orobanche pubescens, Kreta, accessed 7 Oct. 2010).

Italy / Italia: (cf. Bertoloni, 1846: 441; Beck, 1890: 239; Beck, 1930: 184, 186; Uhlich & al., 1995: 125; Domina & Mazzola, 2007: 225; Domina & Arrigoni, 2007: 127).

Sicily / Sicilia: (cf. Lojacono, 1883: 82; Beck, 1890: 239; Greuter & al., 1989: 261; Domina & Mazzola, 2007: 225).

Lebanon / Libano: (cf. Beck, 1930: 185).

Lybia / Libia: (cf. Beck, 1930: 185; Jafri, 1978: 21; Greuter & al., 1989: 261).

Malta: (cf. Beck, 1890: 239; Greuter & al., 1989: 261; Mifsud, Orobanche pubescens in Malta Wild Plants, accessed 7 Oct. 2010).

Palestine [Israel and the Palestinian territories]: (cf. Beck, 1890: 240; Feinbrun-Dothan, 1978: 214; Greuter & al., 1989: 261; Flora of Israel Online, Orobanche pubescens, accessed 7 Oct. 2010).

Russia / Rusia [Caucasus]: (cf. Novopokrovskii & Tzvelev, 1958[2000]: 126; Tzvelev, 1981 [2001]: 454).

Syria / Siria: (cf. Beck, 1930: 185).

Turkey / Turquía: (cf. Reuter, 1847: 27; K. Koch, 1849: 669; Beck, 1930: 184; Gilli, 1982: 15).

Ukraine / Ucrania: (cf. Reuter, 1847: 27; K. Koch, 1849: 669; Beck, 1930: 184; Gilli, 1982: 15).

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References.

Barina, Z. & Pifkó, D. 2008. Additions and amendments to the flora of Albania. Willdenowia 38: 455-464.

Beck, G. 1890. Monographie der Gattung Orobanche. Biblioth. Bot. 19. Theodor Fischer. Kassel.

Beck, G. 1902. Ordo. Orobanchaceae. In Halácsy, E. von. Conspectus florae Graecae II. Lipsiae: Sumptibus Guilelmi Engelmann. [Consp. Fl. Graec. 2, Orobanchaceae: 444-460].

Beck, G. 1922. IX. (CXVII). […] Orobanchaceae novae. Feddes, Repert. 18: 33[465]-40[472].

Beck, G. 1930. IV. 261. Orobanchaceae. In Engler, A. (ed.). Das Pflanzenreich. Regni Vegetabili Conspectus. Wilhelm Engelmann. Leipzig [pp. 1-348].

Bertoloni, A. 1844-1847. Flora Italica sistens plantas in Italia et in insulis circumstantibus sponte nascentes. [...] Vol. VI. Bononiae MDCCCXLIV. Ex typographaeo Haeredum Richardi Masii, sumptibus auctoris. [Orobanche Lin. Gen. p. 321 in Fl. Ital. 6: 419-455].

Boulos, L. 2002 [Dec. 2002]. Flora of Egypt 3 (Verbenaceae - Compositae). Al Hadara Publishing Cairo, Egypt. [Orobanchaceae: 86-96].

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Crouzet, N., Pavon, D. & Michaud, H. 2009 [18 Nov. 2009]. Mise à jour de la liste des plantes vasculaires du département des Bouches-du-Rhône. Bull. Soc. linn. Provence 60: 55-73.

Domina, G. & Arrigoni, P.V. 2007. The genus Orobanche (Orobanchaceae) in Sardinia. Fl. Medit. 17: 115-136.

Domina, G. & Mazzola, P. 2007. The genus Orobanche in Sicily. Taxa described by V. Tineo and M. Lojacono Pojero. Bocconea 21: 223-232.

Domina, G. & Raab-Straube, E. 2010. Orobanche pubescens. Orobanchaceae. In: Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity (accessed 20 July 2010).

Dumont d'Urville, J.S.C. 1822 [Apr. 1822]. Enumeratio plantarum quas in insulis archipelagi aut littoribus Ponti-Euxini, annis 1819 et 1820, collegit atque detexit J. Dumont d'Urville. Parisiis: ex typis D'Hautel, Venitque apud Causette, Bibliopolam (Quai des Grands-Augustins, nº. 35 (bis)). [Enum. Pl. Ponti-Eux.]. ("Reprinted from Mém. Soc. Linn. Paris 1: 255-387. 1822" [cf. TL-2 Online]). [Orobanchaceae: 76].

Feinbrun-Dothan, N. 1978. 98 - Orobanchaceae. Flora Palestina. Part 3. Israel Academy of Sciences and Humanities. Jerusalem. 208-215 pp.

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Gilibert, J.E. 1782. Flora lituanica inchoata. 2 vol. Grodno.

Gilli, A. 1971. Die Orobanchaceen der Türkei. Feddes Repert. 82(6): 381-406.

Gilli, A. 1982. Orobanchaceae (90). In Davis, P.H. (ed.). Flora of Turkey 7: 1-22. Edinburgh.

Greuter, W., Burdet, H.M. & Long, G. 1989. Med-Checklist.. 4: 265-266 [Orobanchaceae]. Genève, Berlin.

Grisebach, A.H.R. 1843-1846. Spicilegium Florae Rumelicae et Bithynicae Exhibens Synopsin Plantarum quas in aest. 1839 legit Auctor A. Grisebach. Braunschweig, 1843-1845 [2 volumes, 6 parts: 1(1), Mar 1843; 1(2/3), Dec¹1843: 2(4), Jul 1844; 2(5/6), Jan 1846]. Spic. Fl. Rumel.

Habashi, C. & Jeanmonod, D. 2008. Orobanchaceae in Compléments au Prodrome de la Flore Corse. Conservatoire et Jardin botaniques, Genève. 126 pp.

Hayek, A. 1924. Zweiter Beitrag zur Kenntnis der Flora von Albanien. Denkschr. Akad. Wiss. Wien, Math. Naturwiss. Kl. 99: 101-224.

Jafri, S.M.H. 1978. Orobanchaceae. In Jafri, S.M.H. & El-Gadi, A. (eds.). Flora of Libya 55: 1-124. Tripoli.

Jordan, A. 1846 [Sept. 1846]. Observations sur plusieurs plantes nouvelles rares ou critiques de la France. Troisième fragment. Septembre 1846. Paris: Maison, libraire, Quai des Augustins, 29; Leipzig: T. O. Weigel, rue du Roi (Lyon. - Imp. de Dumoulin et Ronet, quai St-Antoine, 33.). [Genre Orobanche: 223-227, pl. 9].

Koch, K. 1849. Beiträge zu einer Flora des Orientes. Linnaea 22: 177-338, [Fortsetzung] 597-752. [Orobanchaceae: 658-670].

Lojacono Pojero, M. 1883. Criterii sui caratteri delle Orobanche ed enumerazione delle nuove specie rinvenute in Sicilia. Naturalista Sicil. 2: 80-84, 105-110, 132-136.

Meikle, R.D. 1977-1985. Flora of Cyprus. Kew. 2 vols. [vol. 1 (1977); vol. 2 (1985)].

Mifsud, S. Orobanche pubescens in Malta Wild Plants. www.maltawildplants.com (accessed 7 Oct. 2010).

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Novopokrovskii, I.V. & Tzvelev, N.N. 1958. Orobanchaceae Vent. In Komarov, V. L., (Ch. Ed.). 1934-1964. Flora SSSR 23 : [I]-XXIII, [1]-775. Edit. Akad. Nauk SSSR. Moskva-Leningrad. [Panigrahi, G. (Bot. Ed.). 2000. Bishen Singh Mahendra Pal Singh].

Reynier, A. 1913. L'Orobanche pubescens d'Urv. en Provence; sa validité nominale et spécifique. Bull. Soc Bot. France 60: 325-330.

Reuter, G. 1847. Orobanchaceae. In De Candolle, Alphonse (ed.). Prodromus Systematis naturalis regni vegetabilis... 11: 1-45. [Published on 25 Nov. 1847].

Schneeweiss, G.M., Colwell, A.E., Park, J.M., Jang, C.G. & Stuessy, T.F. 2004. Phylogeny of holoparasitic Orobanche (Orobanchaceae) inferred from nuclear ITS sequences. Mol. Phylogenetics Evol. 30: 465-478. [Abstract / Download].

Schultz, F.W. 1843 [28 Febr. 1843]. Andeutungen zur Kenntnis einiger. Orobanchen Griechenlands. Flora 26: 125-132.

Schultz, F.W. 1845 [21 Dec. 1845]. Bemerkungen über einige Orobanchen und andere Pflanzen. Flora 28(47): 739-741. Regensburg.

Stoyanov, K. 2009. Chorology and critical notes on Orobanche subsect. Minores in Bulgaria. Phytologia Balcanica 15(3): 351-360 [full text article - pdf].

Tzvelev, N.N. 1981 [2001]. Orobanchaceae Vent. In Fedorov, An.A. (ed.). Flora of Russia: The European Part and Bordering Regions 5: 434-461. Nauka Publishers. Leningrad [A.A. Balkema. India]. [Flora Evropeiskoi Chasti SSSR 5].

Uhlich, H., Pusch, J. & Barthel, K.-J. 1995. Die Sommerwurzarten Europas: Gattung Orobanche. Westarp Wissenschaften - Magdeburg.

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