Orobanche_gamosepala_Grupo botánico cantábrico

Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

ISSN: 2386-9666 - Last Updated: 9 October 2023

Accepted Name:

Orobanche gamosepala Reut. in DC., Prodr. 11: 15 (1847) (Piwowarczyk & al. 2019: 54; Piwowarczyk & al. 2023: 42)

Synonyms:

Homotypic synonyms:

≡ Ceratocalyx reuteri Lange in Vidensk. Meddel. Dansk Naturhist. Foren. Kjobenhavn 1863: 52 (1864) ["Obs. ... C. Reuteri nob. (Orobanche gamosepala Reut.)"] (Piwowarczyk & al. 2019: 54; Piwowarczyk & al. 2023: 42)

Heterotypic synonyms:

= Orobanche phoenicea Fisch. ex Beck in Biblioth. Bot. 19: 154 (1890) ["Fischer, in schedis (unpublished name)"], pro syn. (Turland & al. 2018: Art. 36.1b; Piwowarczyk & al. 2019: 54; Piwowarczyk & al. 2023: 42)

= Orobanche phoenicea Fisch. ex Beck in Engl., Pflanzenr. 96: 225 (1930) ["Fisch. in sched. (unpublished name)"], pro syn. (Beck 1890: 154, sub O. gamosepala)

= Orobanche punicea Prescott [in sched., unpublished name]. Ind. loc.: M. Caucasus. Syntypes: K 000061363-64-65 (http://specimens.kew.org/herbarium/K000061363 - http://specimens.kew.org/herbarium/K000061364 - http://specimens.kew.org/herbarium/K000061365) - http://www.kew.org/herbcatimg/665210.jpg - https://www.gbif.org/occurrence/925146862; rev. Ó. Sánchez Pedraja Sep 2016

= Necranthus orobanchoides Gilli in Notes Roy. Bot. Gard. Edinburgh 28: 297 (1968) (Piwowarczyk & al. 2019: 54; Piwowarczyk & al. 2023: 42). Holotype: "Davis & Hedge (D. 29677) / Orobanche / Turkey. Prov. Kars: Yalnizçam dağ [Yalnızçam Dağları / Arsiani Range] between / Yalnizçam and Ardanuç . 2250 m. Rocky / volcanic slopes. Flowers violet-black! / Anthers cream. 16.6.1957". – 2. "Holotype / of Necranthus orobanchoides Gilli / in Notes Roy. Bot. Gard. Edinburgh 28: 197. (1968)". – 3. "Necranthus orobanchoides Gilli / nov. genus, nov, sp. (Typus) / 1966 det. A. Gilli" (https://data.rbge.org.uk/herb/E00130969 - E 00130969.jpg - https://www.gbif.org/occurrence/574799143). Isotype: 1. "Davis & Hedge (D. 29677) / Orobanche / Turkey. Prov. Kars: Yalnizçam dağ [Yalnızçam Dağları / Arsiani Range] between / Yalnizçam and Ardanuç . 2250 m. Rocky / volcanic slopes. Flowers violet-black! / Anthers cream. 16.6.1957". – 2. "Orobanche gamosepala Reut. / Det. E. Teryokhin 12.04.1996" (K 000820831.jpg - http://specimens.kew.org/herbarium/K000820831 - kew.org/herbcatimg/416007.jpg - https://www.gbif.org/occurrence/912618924; rev. Ó. Sánchez Pedraja Sep 2016) (Piwowarczyk & al. 2019: 55)

inicio / top of page

Ind. loc.: Russia: "In pratis humidis prope col. Karass [colony of Karras located 10 kilometers north of Pyatigorsk] et radices montis Matschuka [Mashuk / Maschuka / Машук Mountain, Mount Beshtau, North Caucasus, near Pyatigorsk, Stavropol Krai] Georgiae Caucasicae (h. Hohenacker !) ... (v. s. in h. Hohenacker. )" (Reuter 1847: 15). "Hab. in provinciis caucasicis [in pratis humidis prope coloniam Karass et ad radicem m. Matschuka ! (Hohenack. pl. exs.)]" (Ledebour 1849: 315, sub O. gamosepala). "In pratis humidis prope col. Karass et radices montis Matschuka Georgiae Caucasicae (h. Hohenacker!)" (Boissier 1879: 503).

Typus / Type: Lectoype: 1. "Orobanche gamosepala Reut. / Herb. Dr. a Kühlewein. / pr. Karass & in m. Maschuka [Mashuk / Maschuka / Машук Mountain, Mount Beshtau, North Caucasus, near Pyatigorsk]; majo [m. Kühlewein] / Fl. Caucasica [leg. Hohenacker]". – 2. "Herb. Acad. Petropol. / N.º nov. sp.? / Orobanche gamosepala Reut.! / Fl. Caucas. legit C.A. Meyer 1829. 1830 Hohenacker // Herb. Al de Bunge" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02968131 - P 02968131.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/437107550; rev. Ó. Sánchez Pedraja Sep 2016 [two right-hand specimens (herb. Dr. a Kühlewein)]); isolectotypes: http://coldb.mnhn.fr/catalognumber/mnhn/p/p02968131 - P 02968131.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/437107550 [only one specimen on left side of the sheet (herb. Al de Bunge)]; 1. "Herbarium horti Petropolitani / Orobanche gamosepala Reut. / ! Beck [m. Beck] / Maschuka. Caucasus Hohenacker". – 2. "Typus!" (http://re.herbariumle.ru/01013898 - LE 01013898); 1. "Orobanche / nov. sp.? / Karass". – 2. "I. Novopokrovsky. Notae criticae / Orobanche gamosepala Reut. / specimen a Hohenacker (?) prope stationem / Karass (?) lectum. Cotypus? / 1948.26.XII [?] (vide Reuter in DC. Prodr.)". – 3. "Orobanche gamosepala Reut. / Isotypus! / 1988 III Teste N.Tzvelev" (http://re.herbariumle.ru/01013900 - LE 01013900); 1. "Orobanche / nov. sp.? / Hohenacker / Karass / Herb. Fischer [see label 1 (LE 01013900)]". – 2. "Herb. Fischer / Orobanche / Mai. Maschuka Hohenacker / diff. tubo corolla elongato, ... denticulato". – 3. "N.º 5 / Orobanche Purpurea / wachft in die Ebenen des Berges Beschtau auf Pjatigorsk baden / bluht May [?] / atrorubens / ... / Caucasus / Herb. Fischer / Wilhelms". – 4. "! N.Y. [m. Novopokrovsky?] / O. gamosepala Reuter / det. Dr. G. Beck [m. Beck]". – 5. "Orobanche gamosepala Reut. / 1847, in DC. Podr. 11: 15. / Isotypus! / 1988 III Teste N.Tzvelev" (http://re.herbariumle.ru/01013899 - LE 01013899); 1. "Orobanche n. sp.? / Hohenacker / Karass / ... [pr. Karass & in m. Maschuka] / May". – 2. "I. Novopokrovsky. Notae criticae / Orobanche gamosepala Reut. / Cotypus! / 1948.26.XII". – 3. "Cotypus!" (http://re.herbariumle.ru/01015360 - LE 01015360); 1. "Orobanche gamosepala Reut. / Herb. Dr. a Kühlewein. / pr. Karass & in m. / Maschuka [Mashuk / Maschuka / Машук Mountain, Mount Beshtau, North Caucasus, near Pyatigorsk] leg. / Hohenacker, Majo. [m. Kühlewein] / Fl. Caucasica". – 2. "Identificación confirmada A. Pujadas II 2005" (COI 00056887- herb. Kühlewein - http://coicatalogue.uc.pt/specimen/56887; rev. Ó. Sánchez Pedraja Aug 2018); G (Piwowarczyk & al. 2019: 54).

Designated by: Piwowarczyk & al. in Phytotaxa 386(1): 54 (2019).

Host: Apparently parasitic exclusively on species of Geranium L. (Geraniaceae), but it has also been mentioned as a parasite of Stachys macrantha (K. Koch) Stearn (Lamiaceae).

Geraniaceae Juss. (https://plant.depo.msu.ru/open/public/item/MW0723477 - MW 0723477.jpg - https://www.gbif.org/occurrence/1697840937; https://plant.depo.msu.ru/open/public/item/MW0723482 - MW 0723482.jpg - https://www.gbif.org/occurrence/1697840963; Novopokrovskij & Tzvelev 1958: 88[83], sub O. gamosepala; BioLib.cz 784321, sub O. gamosepala, accessed Aug 2018; Piwowarczyk & al. 2019: 55)

Geranium gracile Ledeb. ex Nordm. [= Geranium trilobum K. Koch, nom. illeg.]. Georgia (Piwowarczyk & al. 2023: 42, sub O. gamosepala).

Geranium gymnocaulon DC. Georgia (Piwowarczyk & al. 2023: 42, sub O. gamosepala); Karachay-Cherkessia (Russia) (https://www.plantarium.ru/page/image/id/526163.html by Дмитрий Бочков [Dmitriy Bochkov] 2017); Krasnodar Krai (Russia) (MW 0723477.jpg).

Geranium ibericum Cav. Georgia (Piwowarczyk & al. 2023: 42, sub O. gamosepala); South Ossetia (Georgia) (W 1941 0003773 - https://www.jacq.org/detail.php?ID=1527225 - https://www.gbif.org/occurrence/2597553399 [n.v.]); Turkey (Rätzel & Uhlich 2004: 193, sub O. gamosepala).

Geranium psilostemon Ledeb. [= Geranium armenum Boiss.; = Geranium backhousianum Regel]. Georgia (Piwowarczyk & al. 2023: 42, sub O. gamosepala).

Geranium sanguineum L. Karachay-Cherkessia (Russia) (https://www.plantarium.ru/page/image/id/189730.html by Сергей Банкетов [Sergey Banquetov] 2013; MW 0723482.jpg).

Geranium sylvaticum L. Georgia (Piwowarczyk & al. 2023: 42, sub O. gamosepala); Adygea (Russia) (Rätzel & Uhlich 2004: 193, sub O. gamosepala); Krasnodar Krai (Russia) ( https://www.inaturalist.org/observations/87244012 - https://www.gbif.org/occurrence/3337376634 by Danila Kurochkin 2021).

Labiatae Juss. [Lamiaceae Martynov, nom. cons., nom. alt. (Turland & al. 2018: Art. 18.5)]

Stachys macrantha (K. Koch) Stearn [≡ Betonica grandiflora Steph. ex Willd.]. Daghestan (Russia) (https://plant.depo.msu.ru/open/public/item/MW0723496 - MW 0723496.jpg - https://www.gbif.org/occurrence/1697841003; Rätzel & Uhlich 2004: 193, sub O. gamosepala).

Comments:

Phylogenetic observation: "putative crucial taxa" (Carlón & al. 2005: 59, sub O. gamosepala).

Taxonomic observations:

"Beck placed the two species in his Orobanche trib. Galeatae Beck in Biblioth. Bot. 19: 134 [152] (1890), nom. invalid. (Turland et al. 2018: Art. 37.6). Syn: ≡ Orobanche [unranked] Galeatae Beck in Halácsy & Braun, Nachtr. Fl. Nieder-Österr.: 125 (1882) (Beck 1890: 152); ≡ Orobanche Grex Galeatae (Beck) Beck in Engl., Pflanzenr. 96: 222 (1930), nom. invalid. Based on genetic studies (Piwowarczyk et al. 2021a), we think that the two species should be considered as a different subgenus (Gamosepalae) within the genus Orobanche. This new subgenus would contain the two sections based on the section (Gamosepalae) and subsection (Arcuatae) published by Teryokhin and placed by him in two different sections (Gamosepalae and Orobanche). The first comprises a single species (O. gamosepala), while the second comprises two (O. colorata and O. anatolica) or one (O. colorata) if we consider them synonyms." (Piwowarczyk & al. 2023: 43-44, sub O. gamosepala).

Orobanche subg. Gamosepalae (Teryokhin) Ó. Sánchez & Piwow., comb. & stat. nov. (Turland et al. 2018: Art. 5A.1 and 6.10) [Orobanche subg. Gamosepalae (Teryokhin) Ó. Sánchez & Piwow. in Phytotaxa 604(1): 43 (2023 [24 Jul 2023])].

Basionym: Orobanche sect. Gamosepalae Teryokhin in Opred. Zarazikhovykh Fl. SSSR (ed. L.Yu. Budantsev): 40 (1993).

Type Name: Orobanche gamosepala Reut. in DC., Prodr. 11: 15 (1847).

—Orobanche sect. Gamosepalae Teryokhin in Opred. Zarazikhovykh Fl. SSSR (ed. L.Yu. Budantsev): 40 (1993).

—Orobanche sect. Arcuatae (Teryokhin) Ó. Sánchez & Piwow., comb. & stat. nov. [Orobanche sect. Arcuatae (Teryokhin) Ó. Sánchez & Piwow. in Phytotaxa 604(1): 44 (2023 [24 Jul 2023])]

Basionym: Orobanche subsect. Arcuatae Teryokhin in Opred. Zarazikhovykh Fl. SSSR (ed. L.Yu. Budantsev): 40 (1993).

Type Name: Orobanche colorata K. Koch. in Linnaea 17: 291 (1843).

Chromosomatic number:

Ic.: a, Beck 1890: t. 2 f. 41, sub O. gamosepala; b, Beck 1930: 224, f. 16 E, sub O. gamosepala; c, Novopokrovsky & Tzvelev 1958: 67[63], t. 5 f. 6, sub O. gamosepala; d, Piwowarczyk & al. 2023: 42 f. 26, sub O. gamosepala.

Distribution: Armenia, Azerbaijan, Georgia, Iran [unconfirmed], Russia, Turkey.

General distribution: "Orobanche gamosepala in Caucasi territoriis indigena a perpaucis locis adhuc cognita est" (Beck 1890: 155, sub O. gamosepala); "General distribution: Probably Armenia-Kurdistan" (Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala); Armenia, Azerbaijan, Georgia, Russia, Turkey (Domina & Raab-Straube 2010, sub O. gamosepala); "General distribution:—the Caucasus, mainly in Ciscaucasia (European Russia), less abundant in Transcaucasia (Georgia, Armenia, Azerbaijan and NE Turkey). Endemic to the Caucasus" (Piwowarczyk & al. 2023: 43, sub O. gamosepala).

inicio / top of page

Specimens examined and literature records.

Armenia / Hayastan / Armenien / Arménie / Әрмәнстан / Армения (Beck 1930: 226, sub O. gamosepala; Novopokrovskii & Tzvelev 1958: 88[83], sub O. gamosepala; Domina & Raab-Straube 2010, sub O. gamosepala; Piwowarczyk & al. 2019: 54-55, figure 37, sub O. gamosepala)

Gegharkunik Province / Гехаркуникская область: "Russisch Armenien: in Bergwiesen bei Akmangan [Mount Armaghan, Vardenis mountain range] nordöstlich von Eriwan [Yerevan] (Seidlitz in Herb. Boissier!)" (Beck 1930: 226, sub O. gamosepala).

Kotayk Province / Котайкская область: (Piwowarczyk & al. 2019: 55, figure 37, sub O. gamosepala).

Syunik Province / Сюникская область: (Piwowarczyk & al. 2019: 55, figure 37, sub O. gamosepala).

inicio / top of page

Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaïdjan [former Azerbaijan Soviet Socialist Republic [Azerbaijan SSR] / Азербайджанская Советская Социалистическая Республика [Азербайджанская CCP] (Boissier 1879: 503, sub O. gamosepala; Beck 1890: 155, sub O. gamosepala; Beck 1930: 226, sub O. gamosepala; Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala; Domina & Raab-Straube 2010, sub O. gamosepala)

Lankaran-Astara Economic Region / Lənkəran-Astara iqtisadi rayonu / Ленкорань-Астаринский экономический район

Lankaran Lowland / Lənkəran ovalığı: "Talysh [Talysh Mountains / Talış dağları / Талыш дағлары]" (Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala).

Karabakh Economic Region / Qarabağ iqtisadi rayonu / Карабахский экономический район (Nagorno-Karabakh Republic / Нагорно-Карабахская Республика) [former Nagorno-Karabakh Autonomous Oblast / Автономная Область Нагорного Карабаха (АОНК)]

1. "Orobanche gamosepala DC. n.l. / Orobanche —— ? / In M. Karagiol raris / sima, tota atropurpurea / Karabagh [Karabakh] Junio 829 Sovitz [O.J. Szovits]". – 2. "Herb. Steven" (http://id.luomus.fi/C.370916 - http://id.luomus.fi/MY.7849674 - https://id.digitarium.fi/api/C.370916/Preview001.jpg - C 370916 - https://www.gbif.org/occurrence/3439931627; rev. Ó. Sánchez Pedraja Oct 2023); "in monte Karagiol (Szov. exs. Nº. 289 !)" (Boissier 1879: 503, sub O. gamosepala); "Indicatur a cl. Boissier: in monte Karagiol (Szov. Exs. nr. 289) prope Achakichad lacum Tabiszchur" (Beck 1890: 155, sub O. gamosepala).

inicio / top of page

Caucasus / Cáucaso / Caucase / Kaukasien / Кавказ (Prescott, loc. class. of O. punicea; Beck 1890: 155, sub O. gamosepala; Beck 1930: 226, sub O. gamosepala; Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala; Domina & Raab-Straube 2010, sub O. gamosepala)

No precise location given: "What I called Orobanche / punicea, is Orobanche / atrorubens m / from M Caucasus" (K 000061363.jpg - http://specimens.kew.org/herbarium/K000061363 - http://www.kew.org/herbcatimg/665210.jpg - https://www.gbif.org/occurrence/925146850; rev. Ó. Sánchez Pedraja Sep 2016); 1. "Orobanche ? / Caucasus / Prescott". – 2. "Orobanche gamosepala Reut DC Prodr. / T. H.G.F.Solms [Hermann Maximilian Carl Ludwig Friedrich zu Solms-Laubach]" (K 000061364.jpg - http://specimens.kew.org/herbarium/K000061364 - http://www.kew.org/herbcatimg/665210.jpg - https://www.gbif.org/occurrence/925146862; rev. Ó. Sánchez Pedraja Sep 2016); "Mr. Prescot. / Orobanche / punicea in M. Caucasus" (K 000061365.jpg - http://specimens.kew.org/herbarium/K000061365 - http://www.kew.org/herbcatimg/665210.jpg - https://www.gbif.org/occurrence/925146763; rev. Ó. Sánchez Pedraja Sep 2016); "Herbarium Universitatis Mosquensis / Orobanche gamosepala Reut. / 1900. Кавказ и Закавказье [Caucasus and Transcaucasia] / [leg.] Ткешелашвили, И.С. [Tkeshelashvili, I.S.]" (https://plant.depo.msu.ru/open/public/item/MW0723476 - MW 0723476.jpg - https://www.gbif.org/occurrence/1697840931; rev. Ó. Sánchez Pedraja Jan 2021); "Herbarium Universitatis Mosquensis (MW) / ... / Orobanche gamosepala Reut. / Кавказ / ..." (https://plant.depo.msu.ru/open/public/item/MW0723475 - MW 0723475.jpg - https://www.gbif.org/occurrence/1697840953; rev. Ó. Sánchez Pedraja Jan 2021).

inicio / top of page

Georgia / Sakartvelo / Georgien / Géorgie / Грузия (former Georgian Soviet Socialist Republic [Georgian SSR] / Грузинская Советская Социалистическая Республика [Грузинская ССР]) (Beck 1890: 155, sub O. gamosepala; Beck 1930: 226, sub O. gamosepala; Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala; Gagnidze 1999; Eristavi & al. 2001: 60, sub O. gamosepala; Gagnidze 2005; Domina & Raab-Straube 2010, sub O. gamosepala; Piwowarczyk & al. 2023: 44, sub O. gamosepala)

Racha-Lechkhumi and Kvemo Svaneti (Piwowarczyk & al. 2023: 44, sub O. gamosepala): "Racha-Lechkhumi" (Eristavi & al. 2001: 60, sub O. gamosepala).

Samegrelo-Zemo Svaneti-Racha-Lechkhumi and Kvemo Svaneti (Piwowarczyk & al. 2023: 44, sub O. gamosepala): "in Swanetia [Svanetia / Svaneti / Suania / Сванетия] (leg. Radde)" (Beck 1890: 155, sub O. gamosepala); "Mingrelien [Samegrelo / Mingrelia / Мегрелия / Мингрелия]: an den Quellen des Flusses Ingur [Enguri River / Ингури] bei 2900 m Seehöhe (Radde!). — Swanetien [Svanetia / Svaneti / Suania / Сванетия] (Radde!), bei Dadiasch [Svaneti Range / Swanetische Kette] und Latpar [Latpari Pass], in 2100 m Seehöhe (Akinfiew!)" (Beck 1930: 226, sub O. gamosepala).

Samegrelo-Zemo Svaneti (Piwowarczyk & al. 2023: 44, sub O. gamosepala): "Svaneti" (Eristavi & al. 2001: 60, sub O. gamosepala); "Georgia, Latpari [Latpari Pass], Svanetie, 13 July 2011" (https://www.biolib.cz/en/taxonimage/id201833/?taxonid=784321 [201833] by Michael Kesl in BioLib.cz; accessed Aug 2018).

Samtskhe–Javakheti (Piwowarczyk & al. 2023: 44, sub O. gamosepala): "Javakheti, Meskheti [Samtskhe / Moschia]" (Eristavi & al. 2001: 60, sub O. gamosepala); "Borjomi-Bakuriani-Akhalkalaki Highway, 2 km south from Bakuriani [Borjomi district, Trialeti Range (Lesser Caucasus Mountains)], 41°44'08.0"N 43°30'33.0"E, 06 June 2005, C. Davidson, det. H. Uhlich, 2012 (https://floraoftheworld.org/taxon/flora/17592186258585 in Flora of the World; accessed Jan 2021).

Autonomous Republic of Adjara / Аджарская Автономная Республика / Adjarie / Adscharien / Аджария [formerly Adjarian ASSR / Adzhar ASSR / Аджарская АCCP] (Gagnidze 1999; Piwowarczyk & al. 2023: 44, sub O. gamosepala)

Republic of Abkhazia / Аҧсны / Abjasia / Республика Абхазия [formerly Abkhaz ASSR / Абхазская АССР] (Beck 1930: 226, sub O. gamosepala; Gagnidze 1999; Eristavi & al. 2001: 60, sub O. gamosepala; Piwowarczyk & al. 2023: 44, sub O. gamosepala)

"Abchasien: in Alpenwiesen an den Quellen des Flusses Seken [Sakeni River, Kodori Range (Greater Caucasus)] (Alboff!)" (Beck 1930: 226, sub O. gamosepala); "Абхазия, Рицинский реликтовый национальный парк [Ritsa Strict Nature Reserve], гора Агура [mountain Agura], урочище Каменная Поляна [Kamennaya Polyana tract, 43°29'48.4"N 40°44'38.0"E], ≈ 2400 м н.у.м., субальпийский луг. 14.07.2017" (http://www.plantarium.ru/page/image/id/553690.html by Марина Скотникова [Marina Skotnikova], 2018 in Plantarium, sub O. gamosepala; accessed Aug 2018 [on Geranium sp.?]).

Republic of South Ossetia – The State of Alania / Tskhinvali Region / Республика Южная Осетия / Государство Алания [formerly South Ossetian Autonomous Oblast / Юго-Осетинская автономная область] (Piwowarczyk & al. 2023: 44, sub O. gamosepala)

"Местность Эрмани [Ermani area - Gora Ermani, 42°29'11.2"N 44°16'10.9"E]. Прав. борт Средне-Эрманского ущелья [Sredne-Erman gorge]. Субальп. луг. На корневучках Geranium ibericum / 1939-07-12, leg. Mozgova, V.D. s.n. / det. M.J.Y. Foley i-1998" (W 1941 0003773 - https://www.jacq.org/detail.php?ID=1527225 - https://w.jacq.org/W19410003773 - https://www.gbif.org/occurrence/2597553399 [n.v.]).

Unknown locations: "prope Plavarzalli Madscha (leg. Frieck)" (Beck 1890: 155, sub O. gamosepala); "in Iberia [Kingdom of Iberia] (Wilhelms)" (Beck 1890: 155, sub O. gamosepala).

inicio / top of page

Iran / Irán / Persia / Persien [unconfirmed]

Ardabil Province-Gilian Province: "Talysh [Talysh Mountains / Talış dağları / Талыш дағлары]" (Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala).

inicio / top of page

Russia / Россия / Rossiya / Rusia / Russie / Russland / 俄罗斯 (Reuter 1847: 15, loc. class. of O. gamosepala; Ledebour 1849: 315, sub O. gamosepala; Boissier 1879: 503, loc. class. of O. gamosepala; Beck 1890: 155, sub O. gamosepala; Beck 1930: 226, sub O. gamosepala; Novopokrovsky & Tzvelev 1958: 88[83], sub O. gamosepala; Domina & Raab-Straube 2010, sub O. gamosepala; Zernov [Зернов] & al. 2015: 379, sub O. gamosepala; Piwowarczyk & al. 2019: 54, sub O. gamosepala)

North Caucasian Federal District / Северо-Кавказский федеральный округ / Severo-Kavkazsky federalny okrug (Zernov [Зернов] & al. 2015: 379, sub O. gamosepala)

Chechen Republic / Чеченская Республика / Chechnya / Чечня: "Чеченская Респ. [42°47'36.6"N 45°54'31.2"E 42.793489, 45.908668, near Nokhchi-Keloi / Нохчи-Келой, Shatoysky District / Шатойский район] 29 May 2022" (https://www.inaturalist.org/observations/119275584 - https://www.gbif.org/occurrence/3859593619 by https://www.inaturalist.org/people/salazar95 [Дауд Арсанукаев / Daud Arsanukaev] in iNaturalist, accessed Oct 2023 [on Geranium sp.?]).

Dagestan / Daghestan / Республика Дагестан (formerly Дагестанская АССР / Dagestan SSSR): "in pratis alpinis montis Antschabala [Bogos Range / Bogosis K’edi / Bogosskiy Khrebet / Богосский Хребет, Mt. Angabala (Gora Addala-Shukhgelmeer / Addala-Shukhgelmeer / Аддала-Шухгельмеэр), "Mons Antschabala. Caucasi", Shamilsky District / Шамильский район] ditionis Tindal [Tlyadal’ / Тлядал /Тлядаль] Caucasi 7000'-8000' (Rupr!)" (Boissier 1879: 503, sub O. gamosepala); "Daghestan: prope Achty [Achty / Ахты, Akhtynsky District / Ахтынский район] (Becker)" (Beck 1890: 155, sub O. gamosepala); "a cl. Ruprecht: in alpinis montis Antschabala [Bogos Range / Bogosskiy Khrebet / Богосский Хребет, Mt. Angabala, "Mons Antschabala. Caucasi"] ditionis Tindal" (Beck 1890: 155, sub O. gamosepala); "Orobanche gamosepala Reut. / Кавказ. Дагестан. / окрестности пос. Кубачи [Kubachi, Dakhadayevsky District / Дахадаевский район]. / Сырои пцг по склоныy N экспозиции / h = 1700 м н.у.м. / ... нa Betonica grandiflora [Stachys macrantha (K. Koch) Stearn] / 24.05.1989 г. / Leg.: В.Д. Бочкин / И.И. Рчсанввич [V.D. Bochkin I.I. Rzsanvic et al.] / . Det. ..." (https://plant.depo.msu.ru/open/public/item/MW0723496 - MW 0723496.jpg - https://www.gbif.org/occurrence/1697841003; rev. Ó. Sánchez Pedraja Aug 2018); "Цунтинский р-н, Респ. [Tsuntinsky District, 42°10'17.5"N 46°01'21.6"E]", 10 jun 2008 (https://www.inaturalist.org/observations/46240379 - https://www.gbif.org/occurrence/2626652656 by https://www.inaturalist.org/people/ramazan_murtazaliev in iNaturalist, accessed Jan 2021 [on Stachys macrantha (K. Koch) Stearn (Betonica grandiflora Steph. ex Willd.)]?); Charodinsky District [Чародинский район, 42°13'17.7"N 46°41'12.0"E], 14 jun 2019 (https://www.inaturalist.org/observations/35793157 - https://www.gbif.org/occurrence/2460070747 by Nadezhda Liksakova in iNaturalist, accessed Jan 2021 [on Stachys sp.?]).

Kabardino-Balkarian Republic / Кабардино-Балкарская Республика: "Herbarium Universitatis Mosquensis (MW) / Orobanche / Кабардино-Балкарская АССР [Kabardino-Balkarian ASSR] / гора Искол [Mount Iskol]. На склоне южной экспозиции. / 17.VII.1979 Суслова [Suslova] (https://plant.depo.msu.ru/open/public/item/MW0723484 - https...MW0723484/img/0.jpg?original=b8ae0c32-d38e-479e-97f3-145a880114aa - MW 0723484.jpg; rev. Ó. Sánchez Pedraja Jan 2021); "Верхушка цветущего растения. Северный Кавказ [North Caucasus], Приэльбрусье [Elbrus region], северный склон г. Чегет [Gora Cheget, 43°14'27.0"N 42°29'57.0"E 43.240833, 42.499167, near Terskol / Терскол (Elbrussky District / Эльбрусский район)], выс. 2800 м н.у.м. Конец июня 2012 г." (https://www.plantarium.ru/page/image/id/158452.html by Александр Лебедев [Alexander Lebedev] 2012; таксон определил(а) Андрей Ковальчук [Andriy Kovalchuk] in Plantarium, sub O. gamosepala, accessed Mar 2022); "Черекский р-н [Chereksky District, 43°03'59.3"N 43°23'33.8"E]", 28 Jun 2020 (https://www.inaturalist.org/observations/60088020 - https://www.gbif.org/occurrence/2868347782 by Nadezhda Liksakova in iNaturalist, accessed Jan 2021); "Эльбрусский район [Elbrussky District, 43°13'42.0"N 42°29'45.9"E 43.228338, 42.496088, near Terskol / Терскол], 8 Jul 2023" (https://www.inaturalist.org/observations/174115548 - https://www.gbif.org/occurrence/4165883426 by https://www.inaturalist.org/people/yulia_necha in iNaturalist, accessed Oct 2023 [on Geranium sp.?]).

Karachay-Cherkess Republic / Карачаево-Черкесская Республика / Karachay-Cherkessia / Карачаево-Черкесия (formerly Karachay-Cherkess Autonomous Oblast / Карачаево-Черкесская АО): "Im Kuban-Gebiete: bei Batalpaschinsk [Batalpashinsk / Cherkessk / Черкесск (Cherkessk Urban Okrug)], im Juni (Lipsky!) ... , im Tale des Flusses Taberda [Valley of the Tebarda River / Reka Teberda / река Теберда] (E.A. Endaurowa!)" (Beck 1930: 226, sub O. gamosepala)" (Beck 1930: 226, sub O. gamosepala); "Herbarium Universitatis Mosquensis / Orobanche gamosepala Reut. / западный Кавказ. Тебердинский заповедник [Teberda Nature Reserve (Карачаевский район / Karachayevsky District)] ... 2300 m / N 291 / 2.VII.1962 / Leg. et det. В. Павлов [V. Pavlov]" (https://plant.depo.msu.ru/open/public/item/MW0723487 - MW 0723487.jpg - https://www.gbif.org/occurrence/1697841027; rev. Ó. Sánchez Pedraja Jan 2021); "Herbarium Universitatis Mosquensis / Orobanche ~~kurdica~~ ~~Boiss.~~ / Ставропольский край. Тебердинский государственный заповедник [Teberda Nature Reserve (Карачаевский район / Karachayevsky District)] / гора Малая Хатипара [Gora Malaya Khatipara]. Восточный / склон ... / 30.VI.1963 [leg.] Двораковская [Dvorakovskaya]". – 2. "Orobanche gamosepala Reut." (https://plant.depo.msu.ru/open/public/item/MW0723492 - MW 0723492.jpg - https://www.gbif.org/occurrence/1697840972; rev. Ó. Sánchez Pedraja Jan 2021 [see MW0669408]); "Herbarium Universitatis Mosquensis / Orobanche gamosepala Reut. / Ставропольский край. Карачаево-Черкесская АО [Karachay-Cherkess Autonomous Oblast]. / Субальпийский луг в долине [Subalpine meadow in the valley of the] реки Гоначхир [river Gonachkhir (Карачаевский район / Karachayevsky District)]. / 16.VII.1967. Собр. Дерягин Г.В. [Deryagin G.V.]" (https://plant.depo.msu.ru/open/public/item/MW0723494 - https...MW0723494/img/0.jpg?original=7383bba4-7b81-4498-b250-0fb5a65ab644 - MW 0723494.jpg - https://www.gbif.org/occurrence/1697840982; rev. Ó. Sánchez Pedraja Jan 2021); Кавказ / Caucasus, Ставропольский край, Тебердинский заповедник [Teberda Nature Reserve (Карачаевский район / Karachayevsky District)] / Домбай гора / Мусса-Ачитара / Dombay Mount / Mussa-Achitara?, 21.VI.1972. Чубатова [Chubatova] (https://plant.depo.msu.ru/open/public/item/MW0723489 - MW 0723489.jpg - https://www.gbif.org/occurrence/1697840985; rev. Ó. Sánchez Pedraja Jan 2021 [see MW0702086]); Ставропольский край [Stavropol Krai], Карачаево-Черкесская авт. обл. [Karachay-Cherkess Autonomous Oblast], 12.VII.1972 (https://plant.depo.msu.ru/open/public/item/MW0723486-2 - MW 0723486-2.jpg - https://www.gbif.org/occurrence/2642854359; rev. Ó. Sánchez Pedraja Jan 2021); "... / Ставропольский край, Карачаево Черкесия / АССР, г. Теберда [Teberda] / ... / 12.07.1973 . [Leg. et det.] Новикова Е.Э. [Novikova E.E.]" (https://plant.depo.msu.ru/open/public/item/MW0723490 - MW 0723490.jpg; rev. Ó. Sánchez Pedraja Jan 2021); Карачаево-Черкесская автономная область [Karachay-Cherkess Autonomous Oblast], Тебердинский заповедник [Teberda Nature Reserve], 5 июля 1984 г. (https://plant.depo.msu.ru/open/public/item/MW0723486-1 - MW 0723486-1.jpg - https://www.gbif.org/occurrence/2642853348; rev. Ó. Sánchez Pedraja Jan 2021); "... / Orobanche gamosepala Reut. / Ставропольский край. / Тебердинский государственный заповедник [Teberda Nature Reserve] / ... Домбайская Поляна [Dombayskaya Polyana, 43°17'26.0"N 41°37'29.0"E (Домбай / Dombay, Karachayevsk / Карачаевск)] ... / 10 июля 1974 г. [leg.] Г. Корнейчук [G. Korneychuk]". – 2. "Orobanche gamosepala Reut." (https://plant.depo.msu.ru/open/public/item/MW0723493 - MW 0723493.jpg - https://www.gbif.org/occurrence/1697840965; rev. Ó. Sánchez Pedraja Jan 2021); "Herbarium Universitatis Mosquensis / Orobanche gamosepala Reut. / ... Карачаево-Черкесская, Тебердинский заповедник [Teberda Nature Reserve] гора Мусса Агмара [Mount Mussa Agmara - Домбай гора / Мусса-Ачитара / Dombay Mount / Mussa-Achitara?] ... 2500 m ... 26.06.1991 г. / Собр. Коджебаш В.Ф. [Kodzhebash V.F.]" (https://plant.depo.msu.ru/open/public/item/MW0723488 - MW 0723488.jpg - https://www.gbif.org/occurrence/1697840968; rev. Ó. Sánchez Pedraja Jan 2021); Orobanche gamosepala Reut. / 11.05.1998. ... / КЧР, Зеленчукский район [Zelenchuksky District], / ... / [Leg.] В. Онипченко [V. Onipchenko] (https://plant.depo.msu.ru/open/public/item/MW0723481 - MW 0723481.jpg - https://www.gbif.org/occurrence/1697840983; rev. Ó. Sánchez Pedraja Jan 2021); "Herbarium Universitatis Mosquensis (MW) / Orobanche gamosepala Reut. / Карачаево-Черкессия, Джегутинский р-н [Dzhegutinsky District], левый берег реки Джегута, холмы между пос. Усть-Джегута [Ust-Dzheguta] и пос. Новая Джегута [Novaya Dzheguta]. На степном склоне. nº 4650 14 мая 2006 г. Leg. А.С. Зернов, Д.К. Текеев [A.S. Zernov, D.K. Tekeyev] Det. А.С. Зернов (https://plant.depo.msu.ru/open/public/item/MW0723483 - MW 0723483.jpg - https://www.gbif.org/occurrence/1899273887; rev. Ó. Sánchez Pedraja Aug 2018); Джегутинский р-н [Dzhegutinsky District], Пастбищный хребет между пос. Усть-Джегута [Ust-Dzheguta] и Новая Джегута [Novaya Dzheguta], средняя часть южного склона. Степной склон. 19 мая 2007 г. Leg. et det. А.С. Зернов [A.S. Zernov] (https://plant.depo.msu.ru/open/public/item/MW0723482 - https...MW0723482/img/0.jpg?original=7767b751-0595-4188-98ef-ae4756e701a9 - MW 0723482.jpg - https://www.gbif.org/occurrence/1697840963; rev. Ó. Sánchez Pedraja Aug 2018 [on Geranium cf. sanguineum]); "Herbarium Universitatis Mosquensis (MW) / Orobanche gamosepala Reut. / Карачаево-Черкесия, Малокарачаевский район [Malokarachayevsky District] ... / Nº 6656 11 мая 2008 г. / Leg. et det. А.С. Зернов [A.S. Zernov]" (https://plant.depo.msu.ru/open/public/item/MW0723485 - MW 0723485.jpg - https://www.gbif.org/occurrence/1697840967; rev. Ó. Sánchez Pedraja Jan 2021); "Республика Карачаево-Черкесия, Домбай [Dombay (Karachayevsk / Карачаевск)], Верховья долины р. Домбай-Ульген [Dombay-Ulgen], в березняке у Чухчурских водопадов [Chukhchur waterfalls], 2100 м н.у.м. 08.07.2011" (http://www.plantarium.ru/page/image/id/98849.html, plantarium.ru/page/image/id/98850 and http://www.plantarium.ru/page/image/id/98851.html by Евгений Комаров [Evgeny Komarov], 2011, [det.] растение определил(а) Холгер Улих [Holger Uhlich], in Plantarium, sub O. gamosepala; accessed Aug 2018 [on Stachys sp.?]); "Расцветающее растение. Карачаево-Черкесская Республика, Карачаевский р-н [Karachayevsky District], окр. пгт Теберда [Teberda], ущелье Азгек [Azgek gorge], окр. оз. Азгек. 14.07.2011" (https://www.plantarium.ru/page/image/id/172268.html by Наталия Жданова [Natalia Zhdanova] 2013; таксон определил(а) Леонид Непоменко [Leonid Nepomenko], in Plantarium, sub O. gamosepala; accessed Mar 2022 [on Geranium sp.?]); "Верхушка цветущего растения. Северный Кавказ, Карачаево-Черкесия, Теберда, ущ. [Teberda gorge] р. Азгек [Azgek river (Reka Teberda / река Теберда)]. 26.07.2011" (https://www.plantarium.ru/page/image/id/126641.html and https://www.plantarium.ru/page/image/id/126642.html by Яна Лисицкая [Yana Lisitskaya] 2012, [det.] растение определил(а) Холгер Улих [Holger Uhlich], in Plantarium, sub O. gamosepala; accessed Jan 2021 [on Geranium cf. gymnocaulon]); "Lagonaki [Caucasian State Nature Biosphere Reserve] og Dombai [Dombay-Ulgen], svært nær grensa til Georgia" (photos Lagonaki 2529 and Lagonaki 2529(2) by Arne Jakobsen in Kaukasus juni 2017 - http://fjellplanter.blogspot.com/, sub O. gamosepala; accessed Aug 2018); "Цветущее растение, паразитирующее на Geranium sanguineum. Карачаево-Черкесия, Теберда, гора Лысая [Teberda, Lysaya Mountain, 43°29'33.0"N 41°42'03.0"E]. 29.05.2013" (https://www.plantarium.ru/page/image/id/189730.html, https://www.plantarium.ru/page/image/id/189733.html and plantarium.ru/page/image/id/189734 by Сергей Банкетов [Sergey Banquetov] 2013 in Plantarium, sub O. gamosepala; accessed Jan 2021); "Цветущее растение (паразитирует на Geranium gymnocaulon). Карачаево-Черкесия, Тебердинский заповедник [Teberda Nature Reserve], хр. Малая Хатипара [Malaya Khatipara ridge, 43°26'46.6"N 41°41'28.3"E], субальпийский луг. 14.07.2017" (https://www.plantarium.ru/page/image/id/526163.html by Дмитрий Бочков [Dmitriy Bochkov] 2017 in Plantarium, sub O. gamosepala; accessed Jul 2021); Карачаевский район [Karachayevsky District, 43°26'44.7"N 41°40'58.8"E, near Teberda / Теберда], 14 Jul 2017 (https://www.inaturalist.org/observations/35579944 - https://www.gbif.org/occurrence/2456561057 by Dmitriy Bochkov in iNaturalist, accessed Jan 2021 [on G. gymnocaulon DC.?]); Карачаево-Черкесская Республика, Усть-Джегутинский [Ust-Dzhegutinsky District] / ... / ... 29 мая 2018 г. (https://plant.depo.msu.ru/open/public/item/MW1008929 - MW 1008929.jpg - https://www.gbif.org/occurrence/3004103750; rev. Ó. Sánchez Pedraja Jan 2021); "Карачаевский район [Karachayevsky District, 43°26'44.7"N 41°40'58.8"E, near Teberda / Теберда], 26 Jul 2018" (https://www.inaturalist.org/observations/36375555 - https://www.gbif.org/occurrence/2596225044 by Dmitriy Bochkov in iNaturalist, accessed Jan 2021 [on G. gymnocaulon DC.?]); "Карачаевский район [Karachayevsky District, 43°26'44.7"N 41°40'58.8"E, near Teberda / Теберда], 25 Jul 2019" (https://www.inaturalist.org/observations/31686436 - https://www.gbif.org/occurrence/2397606772 by Dmitriy Bochkov in iNaturalist, accessed Jan 2021 [on G. gymnocaulon DC.?]); "Краснодарский край [43°37'20.0"N 40°18'05.0"E 43.622222, 40.301389, near Estosadok / Эстосадок (Adlersky City District / Адлерский внутригородской район)], 2 Jul 2022" (https://www.inaturalist.org/observations/125427934 - https://www.gbif.org/occurrence/4076246701 by Сергей Грабчак / Sergey Grabchak in iNaturalist, accessed Oct 2023); Урупский р-н [Urupsky District, 43°37'07.2"N 41°02'30.4"E 43.618678, 41.041778, near Pkhiya / Пхия], 25 Jul 2022 (https://www.inaturalist.org/observations/132112688 - https://www.gbif.org/occurrence/3903277125 by https://www.inaturalist.org/people/itallmas [Екатерина Кропочева / Ekaterina Kropocheva] in iNaturalist, accessed Oct 2023); "Домбай [Dombay (Karachayevsk / Карачаевск), 43°17'21.9"N 41°37'24.6"E 43.289402, 41.623506], 7 Ago 2022" (https://www.inaturalist.org/observations/133914690 - https://www.gbif.org/occurrence/3902374236 by Andrey Efremov in iNaturalist, accessed Oct 2023); "Урупский р-н [Urupsky District, 43°38'44.2"N 40°58'59.2"E 43.645601, 40.983113, near Pkhiya / Пхия], 11 Jul 2023" (https://www.inaturalist.org/observations/174532644 - https://www.gbif.org/occurrence/4165900397 by https://www.inaturalist.org/people/demons1312 [D.S. Shilnikov] in iNaturalist, accessed Oct 2023); "Карачаевский р-н [Karachayevsky District, 43°15'02.2"N 42°14'00.5"E 43.250600, 42.233460 near Aktyube / Актюбе], 23 Jul 2023" (https://www.inaturalist.org/observations/181900513 by Bogdan V. Kryzhatyuk in iNaturalist, accessed Oct 2023 [very illustrative images]).

Republic of North Ossetia-Alania / Республика Северная Осетия-Алания [formerly North Ossetian Autonomous Soviet Socialist Republic / Северо-Осетинская Автономная Советская Социалистическая Республика]: "Herbarium Universitatis Mosquensis (MW) / Северо-Осетинская АССР, бассейн р. Ардон [Ardon river], / южный склон Цейского хребта [Tseysky ridge] (левый борт / Цейского ущелья [Tseysky gorge (Gornyy Kurort Tsey / Горный курорт Цей, near Alagir / Алагир (Alagirsky District / Алагирский район)]) [Субальп. березняки. Высота 1700 м над ур. моря] 14.07.1976. [Leg.] А.М. Амирханов [A.M. Amirhanov]". – 2. "Orobanche gamosepala Reut. / 1977 Teste: Амирханов [Det. Amirhanov]" (https://plant.depo.msu.ru/open/public/item/MW0723495 - MW 0723495.jpg - https://www.gbif.org/occurrence/1697840932; rev. Ó. Sánchez Pedraja Jan 2021); "Алагирский р-н [Alagirsky District, 42°46'58.5"N 43°54'05.4"E 42.782907, 43.901493, near Verkhnii / Tsei Верхний Цей], 30 Jun 2013" (https://www.inaturalist.org/observations/78372956 by https://www.inaturalist.org/people/demons1312 [Шильников Дмитрий Сергеевич / Schilnikow Dmitry Sergejewitsch] in iNaturalist, accessed Oct 2023); "г. Владикавказ [Gorod Vladikavkaz, 42°56'18.8"N 44°36'50.2"E, near Vladikavkaz / Владикавказ], 29 May 2021" (https://www.inaturalist.org/observations/80946138 - https://www.gbif.org/occurrence/3321199482 by https://www.inaturalist.org/people/e_verbitsky in iNaturalist, accessed Mar 2022 [on Geranium sp.?]).

Stavropol Krai / Ставропольский край: 1. "Orobanche gamosepala Reut. / Herb. Dr. a Kühlewein. / pr. Karass [colony of Karras located 10 kilometers north of Pyatigorsk] & in m. Maschuka [Mashuk / Maschuka / Машук Mountain, Mount Beshtau, North Caucasus, near Pyatigorsk]; majo [m. Kühlewein] / Fl. Caucasica [leg. Hohenacker]". – 2. "Herb. Acad. Petropol. / N.º nov. sp.? / Orobanche gamosepala Reut.! / Fl. Caucas. legit C.A. Meyer 1829. 1830 Hohenacker // Herb. Al de Bunge" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02968131 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02968131 - P 02968131.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/437107550 - https://explore.recolnat.org/occurrence/5A64641B287F47F88927215D2A9F6CC5; rev. Ó. Sánchez Pedraja Sep 2016, lectotype of O. gamosepala [two right-hand specimens (herb. Dr. a Kühlewein)] and isolectotype [only one specimen on left side of the sheet (herb. Al de Bunge)]); 1. "Orobanche gamosepala Reut. / Herb. Dr. a Kühlewein. / pr. Karass [colony of Karras located 10 kilometers north of Pyatigorsk] & in m. / Maschuka [Mashuk / Maschuka / Машук Mountain, Mount Beshtau, North Caucasus, near Pyatigorsk] leg. / Hohenacker, Majo. [m. Kühlewein] / Fl. Caucasica". – 2. "Identificación confirmada A. Pujadas II 2005" (COI 00056887- herb. Kühlewein - http://coicatalogue.uc.pt/specimen/56887 - https://www.gbif.org/occurrence/2598814769; rev. Ó. Sánchez Pedraja Aug 2018, isolectotype of O. gamosepala); 1. "Herbarium horti Petropolitani / Orobanche gamosepala Reut. / ! Beck [m. Beck] / Maschuka. Caucasus Hohenacker". – 2. "Typus!" (http://re.herbariumle.ru/01013898 - LE 01013898, isolectotype of O. gamosepala); 1. "Orobanche / nov. sp.? / Karass [colony of Karras located 10 kilometers north of Pyatigorsk]". – 2. "I. Novopokrovsky. Notae criticae / Orobanche gamosepala Reut. / specimen a Hohenacker (?) prope stationem / Karass (?) lectum. Cotypus? / 1948.26.XII [?] (vide Reuter in DC. Prodr.)". – 3. "Orobanche gamosepala Reut. / Isotypus! / 1988 III Teste N.Tzvelev" (http://re.herbariumle.ru/01013900 - LE 01013900, isolectotype of O. gamosepala); 1. "Orobanche / nov. sp.? / Hohenacker / Karass [colony of Karras located 10 kilometers north of Pyatigorsk] / Herb. Fischer [see label 1 (LE 01013900)]". – 2. "Herb. Fischer / Orobanche / Mai. Maschuka Hohenacker / diff. tubo corolla elongato, ... denticulato". – 3. "N.º 5 / Orobanche Purpurea / wachft in die Ebenen des Berges Beschtau auf Pjatigorsk baden / bluht May [?] / atrorubens / ... / Caucasus / Herb. Fischer / Wilhelms". – 4. "! N.Y. [m. Novopokrovsky?] / O. gamosepala Reuter / det. Dr. G. Beck [m. Beck]". – 5. "Orobanche gamosepala Reut. / 1847, in DC. Podr. 11: 15. / Isotypus! / 1988 III Teste N.Tzvelev" (http://re.herbariumle.ru/01013898 - LE 01013899, isolectotype of O. gamosepala); 1. "Orobanche n. sp.? / Hohenacker / Karass / ... [pr. Karass & in m. Maschuka]". – 2. "I. Novopokrovsky. Notae criticae / Orobanche gamosepala Reut. / Cotypus! / 1948.26.XII". – 3. "Cotypus!" (http://re.herbariumle.ru/01015360 - LE 01015360, isolectotype of O. gamosepala); "In pratis humidis prope col. Karass et radices montis Matschuka [Mashuk / Maschuka / Машук Mountain, Mount Beshtau / Бештау, North Caucasus, near Pyatigorsk / Пятигорск] Georgiae Caucasicae (h. Hohenacker !)" (Reuter 1847: 15, loc. class. of O. gamosepala); "Hab. in provinciis caucasicis [in pratis humidis prope coloniam Karass et ad radicem m. Matschuka ! (Hohenack. pl. exs.)]" (Ledebour 1849: 315, sub O. gamosepala); "In pratis humidis prope Karaes [colony of Karras located 10 kilometers north of Pyatigorsk] et radices montis Matschuka Georgiae Caucasicae (h. Hohen !)" (Boissier 1879: 503, loc. class. of O. gamosepala); "Georgia: in pratis humidis prope Karass ad pedem montis Matschuka (Hohenacker loc. class.) (Beck 1890: 155, sub O. gamosepala); "Georgien: in feuchten Wiesen bei Karass am Fuße des Berges Matschuka (loc. class. Hohenacker!)" (Beck 1930: 226, sub O. gamosepala); "Territ. Terek: in monte Besch-tau [Бештау] prope Pjatigorsk (Kolenati et Wilhelms nr. 601)" (Beck 1890: 155, sub O. gamosepala); "Terek-Gebiet: auf dem Beschtau bei Pjatigorsk, in 750-1200 m Seehöhe (Kolenati und Wilhelms n. 604 !, Akinfiew !), um Shelesnowodsk [Zheleznovodsk / Железноводск] (Gerassimow!)" (Beck 1930: 226, sub O. gamosepala); Орджоникидзевский край [Ordzhonikidze Krai, North Caucasus Krai / Северо-Кавказский край], г. Машук [Mashuk] 15.V.1937. Собр. Вернандер Т. [Coll. Vernander T.] (https://plant.depo.msu.ru/open/public/item/MW0723491 - MW 0723491.jpg - https://www.gbif.org/occurrence/1697840948; rev. Ó. Sánchez Pedraja Jan 2021 [see MW0669451]); "Предгорный р-н [43°55'18.5"N 42°46'04.1"E 43.921799, 42.767793, near Kislovodsk / Кисловодск, Predgorny District / Предгорный район], 4 Jun 2017" (https://www.inaturalist.org/observations/78373014 by https://www.inaturalist.org/people/demons1312 [Шильников Дмитрий Сергеевич / Schilnikow Dmitry Sergejewitsch] in iNaturalist, accessed Oct 2023).

Southern Federal District / Южный федеральный округ

Republic of Adygea / Адыгэ Республик / Adyghe Republic / Республика Адыгея: "Orobanche gamosepala Reut. / Кавказский гос. заповедник [Caucasus Nature Reserve], Адыгея, северо-западные отроги горы / Оштен [Mount Oshten], Инструкторская щель, / субальпийский луг. / 18 июля 2004 г. [18.07.2004] N.º 3682 Leg. et det. А.С. Зернов [A.S. Zernov]" (https://plant.depo.msu.ru/open/public/item/MW0723483 - MW 0723483.jpg; rev. Ó. Sánchez Pedraja Aug 2018); "1. Republik Adygeja, NW-Kaukasus, Maikopskij Rajon [Майкопский район], Hauptmassiv des Berges Bolschoi Tchatsch (44°02′33″ N/40°25′20″ E), lichte Staudenflur am Westrand der Alm Poljana Knjažeskaja, Kante zum Kiefern-Birken-Tannen-Wald (licht und staudenreich), Triaskalk, N des Berges bei ca. 1700 m ü.NN, auf Geranium cf. silvaticum [Geranium sylvaticum L.], leg. Rätzel 08.07.2002 Herbar Rätzel 2320, 2321, 2322, det. H. Uhlich 08.2002; – 2. ibidem, Staudenflur (dicht, ca. 1 m hoch) etwa Mitte der Alm Poljana Knjažeskaja, Triaskalk, N des Berges bei ca. 1700 m ü.NN, auf Geranium cf. silvaticum (festgewachsen), nur ein Orobanche-Blütenstand, vergesellschaftet u.a. mit Cirsium Mill., Alchemilla L., Betonica macrantha C.Koch (= B. grandiflora Willd.), Vicia L., Cruciata cf. laevipes Opiz, leg. Fürstenow & Rätzel 08. 07. 2002 Herbar Rätzel 2319 (und Fotos), det. H. Uhlich 08.2002. – 3. NW-Kaukasus, Maikopskij Rajon, subalpin-alpine Hochstaudenflur am N/NE-Abfall des Bolschoi Tchatsch bei >2000 m ü.NN über Triaskalk, leg. Ahrns 28.06.1997 HAL 82695! sub O. caryophyllacea, det. C. Ahrns 23.10.1997, rev. H. Uhlich 11.09.2000" (Rätzel & Uhlich, 2004: 193-194, sub O. gamosepala); Maykopsky District / Майкопский район (https://www.inaturalist.org/observations/8404618 - https://www.gbif.org/occurrence/1677300551 by heeni in iNaturalist, accessed Aug 2018); "Адыгея, Кавказский биосферный заповедник, долина р. Киша [Caucasus Nature Reserve, valley Kisha river (43°46'00.0"N 40°16'00.0"E, Maykopsky District / Майкопский район)], субальпика, 2040 м н.у.м. 01.07.2014" (https://www.plantarium.ru/page/image/id/248572.html et plantarium.ru/page/image/id/248573 by Елена Алешичева [Elena Aleshicheva], 2014, растение определил(а) Владислав Григоренко [Vladislav Grigorenko] in Plantarium, sub O. gamosepala; accessed Aug 2018 [on Geranium sp.]); "Адыгея, Кавказский биосферный заповедник [Caucasus Nature Reserve], под северо-западной стеной горы Фишт, ≈ 1200 м н.у.м., на границе леса и луга. 02.07.2017" (http://www.plantarium.ru/page/image/id/520495.html et plantarium.ru/page/image/id/520496 by Татьяна Гайдаш [Tatiana Gaydash], 2017 in Plantarium, sub O. gamosepala; accessed Aug 2018).

No precise location given: "Soviet Central Asia: All regions besides Talysh" (Novopokrovskii & Tzvelev 1958: 88[83], sub O. gamosepala).

inicio / top of page

Turkey / Türkiye / Turquía / Турция (Gilli 1982: 3, loc. class. of Necranthus orobanchoides; Domina & Raab-Straube 2010, sub O. gamosepala; Piwowarczyk & al. 2019: 55, figure 37, sub O. gamosepala).

Eastern Anatolia Region / Doğu Anadolu Bölgesi

Kars Province / Kars ili: 1. "Davis & Hedge (D. 29677) / Orobanche / Turkey. Prov. Kars: Yalnizçam dağ [Yalnızçam Dağları / Arsiani Range] between / Yalnizçam and Ardanuç . 2250 m. Rocky / volcanic slopes. Flowers violet-black! / Anthers cream. 16.6.1957". – 2. "Holotype / of Necranthus orobanchoides Gilli / in Notes Roy. Bot. Gard. Edinburgh 28: 197. (1968)". – 3. "Necranthus orobanchoides Gilli / nov. genus, nov, sp. (Typus) / 1966 det. A. Gilli" (https://data.rbge.org.uk/herb/E00130969 - E 00130969.jpg - https://www.gbif.org/occurrence/574799143, holotype of Necranthus orobanchoides); 1. "Davis & Hedge (D. 29677) / Orobanche / Turkey. Prov. Kars: Yalnizçam dağ [Yalnızçam Dağları / Arsiani Range] between / Yalnizçam and Ardanuç . 2250 m. Rocky / volcanic slopes. Flowers violet-black! / Anthers cream. 16.6.1957". – 2. "Orobanche gamosepala Reut. / Det. E. Teryokhin 12.04.1996" (K 000820831.jpg - http://specimens.kew.org/herbarium/K000820831 - https://www.kew.org/herbcatimg/416007.jpg - https://www.gbif.org/occurrence/912618924; rev. Ó. Sánchez Pedraja Sep 2016, isotype of Necranthus orobanchoides); "A9 Kars: Yalnizçam dağ [Yalnızçam Dağları / Arsiani Range] between Yalnizçam and Ardanuç, 2250 m, rocky volcanic slopes, 16 vi 1957, I.C. Hedge, D. 29677 (holo. E!)" (Gilli 1982: 3, holotype of Necranthus orobanchoides); "near Sarikamish, 5 July 1911, M. Ivashenko (ERE)" (Piwowarczyk & al. 2019: 55, sub O. gamosepala).

East Black Sea Region / Doğu Karadeniz Bölgesi

Artvin Province / Artvin ili: "Provinz Artvin, Kackar-Gebirge [Kaçkar Mountains], Talschluss südlich von Yaylalar (Hevek), alpine Matten über Silikatgestein, 3000 m ü.NN, an Geranium ibericum, leg. Raus 25.07.2001 B s.n., det. H. Uhlich 25.08.2001" (Rätzel & Uhlich 2004: 193, sub O. gamosepala).

inicio / top of page

References.

Beck, G. 1890. Monographie der Gattung Orobanche. Biblioth. Bot. 19. Theodor Fischer. Kassel. URL: https://bibdigital.rjb.csic.es/idurl/1/10221.

Beck, G. 1930 [30 Sep 1930]. IV. 261. Orobanchaceae Lindl. In: Engler, A. (ed.). Das Pflanzenreich. Regni Vegetabili Conspectus. Wilhelm Engelmann. Leipzig, pp. 1-348. URL: https://www.biodiversitylibrary.org/item/71652.

Boissier, P.É. 1879 [Apr-May 1879]. Flora orientalis [...] Volumen quartum. Fasciculus secundus. Genevae et Basileae. URL: https://bibdigital.rjb.csic.es/idurl/1/10750. [Ord. LXXXIX. Orobanchaceae: 492-518].

Carlón, L., Gómez Casares, G., Laínz, M., Moreno Moral, G., Sánchez Pedraja, Ó. & Schneeweiss, G.M. 2005 [Jun 2005]. Más, a propósito de algunas Orobanche L. y Phelipanche Pomel (Orobanchaceae) del oeste del Paleártico [More on some Orobanche L. and Phelipanche Pomel (Orobanchaceae) from the western Palearctic]. Documentos Jard. Bot. Atlántico (Gijón) 3: 1-71 (in Spanish, plus an Appendix in English).

Domina, G. & Raab-Straube, E. von. 2010. Orobanchaceae. Orobanche gamosepala. In: Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity. URL: http://ww2.bgbm.org/EuroPlusMed/PTaxonDetail.asp?NameId=69633&PTRefFk=7500000 (accessed Aug 2018).

E - RBGE (The Royal Botanic Garden Edinburgh). Herbarium. Databases. URL: https://www.rbge.org.uk/ - http://www.rbge.org.uk/databases/herbarium-catalogue - http://data.rbge.org.uk/herb/E00130969 [browser link]... - https://data.rbge.org.uk/search/herbarium/?specimen_num=153216&cfg=zoom.cfg&filename=E00130969.zip [image link]... (accessed Feb 2015).

Eristavi, M., Shulkina, T., Sikhuralidze, S. & Asieshvili, L. 2001 [s.dat.]. Rare, Endangered and Vulnerable Plants of the Republic of Georgia. URL: http://www.mobot.org/MOBOT/research/georgia/checklist.pdf (accessed Feb 2021).

Gagnidze, R. 1999. Flora Gruzii (Ed. 2) 12. Tbilisi.

Gagnidze, R. 2005. Vascular Plants of Georgia. A Nomenclatural Checklist. Sak’art’velos Mec’nierebat’a Akad, Tbilisi, pp. 1-248.

GBIF (The Global Biodiversity Information Facility). GBIF Online Resource Centre. Search. Orobanchaceae. URL: https://www.gbif.org/ - https://www.gbif.org/species/3733399 (accessed Jul 2017).

Gilli, A. 1968. Necranthus: a new genus of Orobanchaceae from Turkey. Notes. Bot. Gard. Edinburgh 28(3): 297-298.

Gilli, A. 1982. Orobanchaceae (90). In: Davis, P.H. (ed.). Flora of Turkey 7: 1-22. Edinburgh.

iNaturalist. URL: https://www.inaturalist.org/ - https://www.inaturalist.org/taxa/ - https://www.inaturalist.org/taxa/566663-Orobanche-gamosepala - https://www.inaturalist.org/observations/8404618 (accessed Aug 2018).

K (Kew). Royal Botanic Gardens. 2002. Kew Databases - Herbarium catalogue - ePIC [electronic Plant Information Centre] - [searchpage IK]. URL: http://www.kew.org/epic/ - http://specimens.kew.org/herbarium/K000820831 [permalink]... - https://www.kew.org/herbcatimg/416007.jpg [image link]... (accessed Jun 2015).

Lange. J.M.C. 1863-1864 [1864]. Pugillus plantarum imprimis hispanicarum, quas in itinere 1851-52 legit 3. Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn 1863(1-4): 1-58. Kjøbenhavn. Bianco Luno Bogtrykkeri ved. F. S. Muhle. 1864. [XLI Orobancheae, pp. 50-52]. URL: https://bibdigital.rjb.csic.es/idurl/1/11975 - https://www.biodiversitylibrary.org/item/110778.

LE. Vascular Plants Herbarium of the Komarov Botanical Institute RAS — Herbarium LE. Virtual herbarium of Komarov Botanical Institute RAS. URL: http://en.herbariumle.ru/ - http://re.herbariumle.ru/01013898 [permalink - Record creation: 2019-11-26, Ivan Tatanov, PhotoScan D1]... - http://en.herbariumle.ru/?t=occ&id=12150&rid=image_0025581 [image link]... (accessed Feb 2021).

Ledebour, C.F. 1849 [Jun 1849]. Flora rossica 3(1). Stuttgartiae, Sumptibus Librariae E. Schweizerbart, 1847-1849. [Flora Rossica; sive, Enumeratio plantarum in totius Imperii Rossici provinciis Europaeis, Asiaticis et Americanis hucusque observatarum, auctore Carolo Friderico a Ledebour. Volumen III. Pars I. Corolliflorae: Ordo XCIII. Orobanchaceae. Lindl.: 310-328. 1847-1849 (Jun 1849)]. DOI: https://doi.org/10.5962/bhl.title.6606.

MNHN (Muséum National d'Histoire Naturelle), Paris (France). Vascular plants (P). Orobanchaceae. URL: https://science.mnhn.fr/all/search - https://science.mnhn.fr/institution/mnhn/collection/p/item/search - http://coldb.mnhn.fr/catalognumber/mnhn/p/p04419774 [permalink]... - https://science.mnhn.fr/institution/mnhn/collection/p/item/p04419774 [browser link]... - http://mediaphoto.mnhn.fr/media/1441359262314xKYKbF8HNXZfFbfl [image link]... - http://sonneratphoto.mnhn.fr/2011/12/27/3/P04419774.jpg [former image link]... (accessed Nov 2011).

Novopokrovsky, I.V. & Tzvelev, N.N. 1958. Orobanchaceae. In: Shishkin, B.K. (ed.). Flora Unionis Republicarum Socialisticarum Sovieticarum (Flora URSS), vol. 23. Institutum Botanicum nomine V. L. Komarovii Academiae Scientiarum URSS, Mosqua and Leningrad, pp. 685-687. (Novopokrovskii, I.V. & Tzvelev [Cvelev], N.N. 1958. Orobanchaceae Vent. In: Komarov, V. L., (Ch. Ed.). 1934-1964. Flora SSSR 23: 17-111. Edit. Akad. Nauk SSSR. Moskva-Leningrad. [Pandit, V. (Translator & Editor). 1999. Amerind Publishing Co. Pvt. Ltd., New Delhi. Translated and published under an agreement with the Smithsonian Institution Libraries, Washington, DC, by Amerind Publishing Co., 66 Janpath, New Delhi 110 001. 2000] et [Panigrahi, G. (Bot. Ed.). 2000. Bishen Singh Mahendra Pal Singh]). URL: http://www.biodiversitylibrary.org/page/29987536.

Piwowarczyk, R., Sánchez Pedraja, Ó., Moreno Moral, G., Fayvush, G., Zakaryan N., Kartashyan N. & Aleksanyan, A. 2019 [3 Jan 2019]. Holoparasitic Orobanchaceae (Cistanche, Diphelypaea, Orobanche, Phelipanche) in Armenia: distribution, habitats, host range and taxonomic problems. Phytotaxa 386(1): 001-106. DOI: https://doi.org/10.11646/phytotaxa.386.1.1.

Piwowarczyk, R., Schneider, A.C., Góralski, G., Kwolek, D., Denysenko-Bennett, M., Burda, A., Ruraż, K., Joachimiak, A.J. & Sánchez Pedraja, Ó. 2021 [12 Mar 2021]. Phylogeny and historical biogeography analysis support Caucasian and Mediterranean centres of origin of key holoparasitic Orobancheae (Orobanchaceae) lineages. PhytoKeys 174(3):165-194. DOI: https://doi.org/10.3897/phytokeys.174.62524. URL: https://phytokeys.pensoft.net/article/62524 - https://www.researchgate.net/publication/350021224.

Piwowarczyk, R., Sánchez Pedraja, Ó., Khutsishvili, M. & Kharazishvili, D. 2023 [24 Jul 2023]. Holoparasitic Orobanchaceae in Georgia (Caucasus): taxonomic revision, diversity, distribution, habitats and host range. Phytotaxa 604(1): 1-103. DOI: https://doi.org/10.11646/phytotaxa.604.1.1. URL: https://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.604.1.1 - https://www.researchgate.net/publication/372570616.

Plantarium. Плантариум: открытый онлайн атлас-определитель растений и лишайников России и сопредельных стран [Plantarium: open on-line atlas and key to plants and lichens of Russia and neighbouring countries]. 2007-2020. Определитель растений on-line [The determinant of plants on-line]. Поиск таксонов [Search taxa]. Scrophulariaceae. Cistanche. Orobanche. Phelipanche. Phelypaea. URL: http://www.plantarium.ru/ - https://www.plantarium.ru/page/view/item/25917.html (accessed Dec 2014).

Rätzel, S. & Uhlich, H. 2004 [May 2004]. Orobanche benkertii sp. nova (Orobanchaceae Vent.) und weitere Orobanche-Sippen aus dem Nordwest-Kaukasus [Orobanche benkertii sp. nova (Orobanchaceae Vent.) and further Orobanche species from the Northwest Caucasus mountains]. Feddes Repert. 115: 189-211. DOI: https://doi.org/10.1002/fedr.200311036.

Reuter, G. 1847 [25 Nov 1847]. Orobanchaceae. In: De Candolle, A. (ed.). Prodromus Systematis naturalis regni vegetabilis... 11: 1-45. Addenda et Corrigenda: 717-720. Parisiis: Sumptibus Victoris Masson. URL: https://www.biodiversitylibrary.org/page/160970. (TL-2: 446 n. 999).

Seregin, A.P. (ed.). 2017. Specimen MW0723482... from the collection "Moscow University Herbarium" // Depository of Live Systems (branch "Plants"): Electronic resource. – Moscow State University, Moscow. URL: https://plant.depo.msu.ru/open/public/item/MW0723482... (accessed Jun 2018).

Zernov, A.C., Alekseev, Yu.E. & Onipchenko, V.G. [Зернов, А.С., Алексеев, Ю.E. & Онипченко, В.Г.]. 2015. Определитель сосудистых растений Карачаево-Черкесской Республики [The determinant of vascular plants of the Karachay-Cherkess Republic]. Москва.

inicio / top of page

Home page (Página principal) | Index of Orobanchaceae | Contact us