Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors


Accepted Name:

Orobanche cumana Wallr., Orobanches Gen. Diask.: 58 (1825)

Synonyms:

Orobanche arenaria M. Bieb., Fl. Taur.-Caucas. 2: 83 (1808), non Borkh. in Neues Mag. Bot. 1: 6 (1794) (cf. Reichenbach, 1829: 45; Steudel, 1841: 30; Tzvelev, N.N. 1981 [2001]: 451 "The type of O. arenaria Bieb. ... is the type of O. cumana Wallr.)

Orobanche comana [orthographic variant (cf. Schultz, 1830: 495; K. Koch, 1849: 670)]

Orobanche cernua subsp. cumana (Wallr.) Soó in Feddes Repert. 83: 187 (1972)

Orobanche cernua var. cumana (Wallr.) Beck in Engl., Pflanzenr., Orobanchac.: 128 (1930) ["Var. δ cumana"]

Orobanche cernua f. cumana (Wallr.) Beck in Biblioth. Bot. 19: 143 t. 2 f. 33(3) (1890) ["Formae ... 4. (Orobanche cumana Wallroth")]

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= Orobanche bicolor C.A. Mey. in Ledeb., Fl. Altaica 2: 458 (1830). Syn.: Orobanche cernua var. bicolor (C.A. Mey.) Reut. in DC., Prodr. 11: 32 (1847); = Orobanche cumana Wallr. (1825) (sec. Beck, 1930: 128; Novopokrovskii & Tzvelev, 1958: 95). Ind. loc.: "Hab. in deserto soongoro - kirghisico occidentali hinc inde in locis argillosis atque in locis sabulosis ad ripam rivuli Tschaganka versus radicem montis Tschingistau [Eastern Kazakhstan / Kazakstan]" (cf. C.F. Ledebour, Icon. Pl. 4: 25. 1833; "Sibiriae altaicae deserto soongoro - kirghisico! (Fl. Alt., Karel. et Kiril)"; C.F. Ledebour, Fl. Ross. 3: 321. 1841-1853). Ic.: C.F. Ledebour, Icon. Pl. 4: t. 390 (1833). Obs. Collections of S.G.Karelin from Middle Asia (LE)

= Orobanche glaucantha Trautv. in Trudy Imp. S.-Peterburgsk. Bot. Sada 1: 26 (1871) [Acta Horti Petropol. 1: 26 (1871)] (sec. Beck, 1890: 143; Beck, 1930: 128; view IPNI entry). In loc.: "Sibiria orienttali prope Ust Balei [Irkutsk Oblast, Russia]"

= Orobanche sabulosa Steven ex Beck in Biblioth. Bot. 19: 143 (1890) ["in schedulis!"], nom. illeg. Ind. loc.: "provinciis caucasicis [pr. Stawropol (Stavropol / Stauropolis, Stavropol Krai, Russia)] (C.A. Meyer), ad fl. Terek [Terek River, Caucasus]! (Steven pl. exs.)]..." (cf. C.F. Ledebour, 1841-1853: 322, sub O. cumana)

= Orobanche sarmatica Kotov, Fl. URSR 10: 454, f. 8 (1961) (Tzvelev, 1981: 451; view IPNI entry). "near Yevpatoriya, Crimea [Crimea / Krym], UA, (KW)" (cf. Schlauer & al., 2003)

= Orobanche cernua subsp. rajahmundrica Teryokhin in Adv. Paras. Pl. Res.: 246 (1996) (sec. Schlauer & al., 2003) [Weed Broomrapes: 15 (1997), without type. (sec. view IPNI entry)] [in Novost Sist. Vysš. Rast. 33: 210 (2001) (sec. view IPNI entry ; Domina & Raab-Straube, 2010)]. In loc.: "Central Tobacco Research Institute, Rajahmundry, Andhra Pradesh, India, C.A. Raju s.n. (LE)" (cf. Schlauer & al., 2003)

= Orobanche cernua subsp. parviflora Kotov, Sov. Bot. 4: 90 (1935) (cf. Tzvelev, 1981: 451)

= Orobanche cernua var. bicolor (C.A. Mey.) Reut. in DC., Prodr. 11: 32 (1847) ["ß. bicolor"]. Syn.: Orobanche bicolor C.A. Mey. in Ledeb., Fl. Altaica 2: 458 (1830) (cf. C.F. Ledebour, Fl. Ross. 3: 321. 1841-1853)

= Orobanche cernua var. cumana f. densior Beck in Engl., Pflanzenr., Orobanchac.: 128 (1930) ["Forma 4. densior"]. In loc.: "Hinc inde cum varietate"

= Orobanche cernua var. cumana f. holosepala Beck in Engl., Pflanzenr., Orobanchac.: 128 (1930). In loc.: "Shungurei: bei Alakulj (Bienert)" [Songaria / Soogaria / Zhungaria / Dzungaria, Kazakhstan / Kazakstan]"

= Orobanche cernua var. cumana f. taurica Beck in Engl., Pflanzenr., Orobanchac.: 128 (1930) ["Forma 7. taurica"]. In loc.: "In der Krim [Crimea / Krym] bei Oreanda (V. Golde! in Herb. Petersburg. [LE] unter O. alba)"

= Orobanche cernua var. cumana f. tenuisepala (Beck) Beck in Engl., Pflanzenr., Orobanchac.: 128 (1930) ["Forma 6. tenuisepala"]. Syn.: Orobanche cernua f. tenuisepala Beck in Biblioth. Bot. 19: 143 (1890)

= Orobanche cernua f. tenuisepala Beck in Biblioth. Bot. 19: 143 (1890) ["Formae ... 5. (Orobanche) tenuisepala"]. In loc.: "Turkestania: ... in planitie Kickdan ad montes Gasi Machik et prope Gomut [Tajikistan / Tayikistán] (f. 5 leg. Regel)" (cf. Beck, 1890: 145)

= Orobanche cernua var. cumana f. valens Beck in Engl., Pflanzenr., Orobanchac.: 128 (1930) ["Forma 4*. valens"]. In loc.: "Mesopotamien: bei Tell Halaf in der Steppe bei Haus, im Mai (Kohkl! in der Expedition Frh. v. Oppenheim) [Tell Halaf, Al Hasakah, Syria]"

Orobanche cernua auct. pl., non Orobanche cernua L. in Loefl., Iter Hispan.: 152 (1758) [e.g., "O. cérnua Pall. ind. taur." (cf. Don, 1838: 631; Steudel, 1841: 230, "sec. Meyer")]

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Ind. loc.: «Sands in the lower reaches of the Volga and on the Kuma rivers ("In arena mobili deserti astracanici et cumani"). ... "O. arenaria M. In arena mobili ad Wolgam versus Astrachan lecta, 1803"» [Province of Astrakhan / Astrakhan Oblast, Russia] (sec. Tzvelev in Flora of Russia: 451. 1981).

Typus / Type: LE. «The type of O. arenaria Bieb., preserved in Leningrad [LE] with the label "O. arenaria M. In arena mobili ad Wolgam versus Astrachan lecta, 1803," is the type of O. cumana Wallr.» (sec. Tzvelev in Flora of Russia: 451. 1981).

Designated by: Tzvelev in Flora of Russia: 451 (1981).

Host: Parasitic on Compositae (Asteraceae) [e.g., Artemisia sp.] and Solanaceae species [e.g., Nicotiana sp.]. In Europe and other parts of the world is parasitic on various cultivated plants (e.g., Helianthus annus L.) (cf. Annotated Checklist of Host Plants of Orobanchaceae, accessed 24 August 2009).

Comments: Orobanche cumana is a polymorphous species. It is provisionally considered by us to be an independent species very closely allied to Orobanche cernua, although the differences between these two species can not always be easily expressed (cf. Novopokrovskii & Tzvelev, 1958: 95-97; Tzvelev, 1981: 451; Pujadas & Velasco, 2000). We can easily distinguish these taxa when comparing typical specimens (e.g., when comparing specimens of Orobanche cumana growing on cultivated plants) but this is not easy when the specimens are less typical. On the other hand, we believe the indumentum staminal (glabrous / hairy) are of no value in differentiating the two taxa (e.g., O. cernua from loc. class. has hairy stamens) as some authors think (cf. Novopokrovskii & Tzvelev, 1958: 95-96; Tzvelev, 1981: 450-451).

"The form of [O. cernua] subsp. cumana parasitizing sunflower ... result of anthropogenic activity, which, in any case, cannot be considered as type subspecies cumana." (cf. Tzvelev, 1981: 451).

"According to Morozov (1947), the first reports of broomrape in sunflower came from Saratov Oblast in Russia and date back to the 1890s" (cf. Škorić & al. 2010: 64).

Documentos Jard. Bot. Atlántico (Gijón) 3: 61 f. 4 ["Phylogenetic relationships"], 64 (2005).

Documentos Jard. Bot. Atlántico (Gijón) 6: 13 f. 2 [phylogenetic relationships] (2008).

Chromosomatic number: n = 19 (cf. T. Palomeque & A. Sañudo, 1981, sub O. cernua [Granada]), 2n = 38 (G.M. Schneeweiss & al., 2004, sub var. cumana); 2n = 24 (cf. Žukov, 1939, sub O. cumana; Tzvelev, 1981: 451, sub subsp. cumana).

Ic.: a, Reichenbach, 1829: 45, t. 691 f. 926 ["Caucaso: Blume, ex herb. cl.Trevirani"] f. 927 ["Specimen Wallrothianum"]; b, Beck in Biblioth. Bot. 19: t. 2 f. 33(3) (1890); c, Beck, 1930: 113 f. 13; d, Ledebour, Ic. Pl. Fl. Ross.: t. 390 (1833, sub O. bicolor C.A. Mey.); e, habit and corolla ["with host plant Artemisia sp." (cf. Tzvelev in Flora of Russia: 437 t. 41 f. 4. 1981; Novopokrovskii & Tzvelev. Flora SSSR 23: t. 5 f. 3. 1958; Beck in Biblioth. Bot. 19: t. 2 f. 33(3). 1890; Moscow State University Herbarium (MW); f, sheet (Almendros, Cuenca); g, habit (sub var. cumana on its host Artemisia meyeriana (Asteraceae); Georgia, May 2001 by G. Schneeweiss. SIUC, accessed 4 Mar. 2006); h, habit (sub O. cernua. Parasitizing sunflower. Bulgaria, by L.J. Musselman. SIUC, accessed 4 Mar. 2006).

Distribution: Eurasian distribution (see Domina & Raab-Straube, 2010, sub Orobanche cumana, accessed 20 July 2010). Area and zones of severity of Sunflower Broomrape by Yakutkin & Budrevskaya, 2006. Iberian distribution.

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Specimens examined and literature records.

Afghanistan / Afganistán: Afghanistan (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Algeria / Argelia: "Also in northern Africa" (cf. Kreutz, 1995: 88).

Armenia: "Caucas. [Caucasus]" (Steudel, 1841: 30, sub O. cumana); "Caucasi" (cf. Reuter, 1847: 32, sub O. cumana); t. 691 f. 926 ["Caucaso: Blume, ex herb. cl.Trevirani"] (cf. Reichenbach, 1829: 45); "in Caucaso a Blume lectae herbarii Treviraniani" (cf. Reichenbach, 1862: 107).

Azerbaijan / Azerbaiyán / Azərbaycan: "In Tatarisch [Tatarstan, Russia]-Grusien [Georgia] und Schirwan [Shirvan, Azerbaijan] auf Beifuss [Artemisia], 500-800' hoch." (cf. K. Koch, 1849: 670); "Territorium Caucasi: f. 4 (! Helenendorf [Goygol / Xanlar / Chanlar / Elenendorf] prope Jelisawetpol [Ganja / Gəncə / Yelizavetpol]...)" (cf. Beck, 1890: 145, sub O. cernua f. cumana).

Australia [unconfirmed]: Australia (cf. Carter & Cooke, 1994: 586 "potential agricultural weeds in southern Australia"; Kreutz, 1995: 88; Škorić & al. 2010: 64).

Bulgaria: "Bulgarien: bei Varna (nach Davidoff, Prinoss za izont. fl. Varn. XXI [1905]), in der Stara Plan, bei Dragoman in 800 m Seehöhe (F.A. Novak!)." (cf. Beck, 1930: 123, sub O. cernua var. cumana); A new physiological race of broomrape ... observed in Varna ... wild host (Artemisia maritima L. [A. maritima auct. non L. = A. santonicum L.])... (cf. Petrov, 1968; Encheva & Shindrova, 1994: 621); Bulgaria (cf. Petrov, 1970; Greuter & al., 1989: 258, sub O. cernua subsp. cumana; Encheva & Shindrova, 1994: 619, "From its first appearance in 1935 ["in the norteast of Bulgaria"] the parasite has been the constant companion of sunflower ... According to Knyazkov (1950), the severest attacks were recorded in the northeast of Bulgaria (the regions of Varna, Shumen, Provadia"; Shindrova, 1994: 145 f. 1; Gagne, 1998; Shindrova, 2006); "Parasitizing sunflower in Bulgaria" (cf. L.J. Musselman, sub O. cernua. SIUC, accessed 4 Mar. 2006).

China: "Ku pei Ku (f. 4 leg. Wawra)" (cf. Beck, 1890: 145, sub O. cernua f. cumana; Beck, 1930: 125, sub O. cernua var. cumana; Tzvelev, 1981: 451, sub O. cernua subsp. cumana), "Dzhungaria-Kashgaria" [Dzungaria - Kashgar, Xinjiang Uyghur Autonomous Region]; "Gansu, Hebei, W Jilin, Nei Mongol, Qinghai, Shaanxi, Shanxi, Xinjiang" (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana); China (cf. Fernández Escobar & al., 2009).

France / Francia [unconfirmed]: "northward to southern France" (cf. Kreutz, 1995: 88).

Georgia: "Caucas. [Caucasus]" (Steudel, 1841: 30, sub O. cumana); "Caucasi" (cf. Reuter, 1847: 32, sub O. cumana); "ad fl. Terek [Terek River]! (Steven pl. exs.)]..." (cf. C.F. Ledebour, 1841-1853: 322, sub O. cumana); "In Tatarisch [Tatarstan, Russia]-Grusien [Georgia] und Schirwan [Shirvan, Azerbaijan] auf Beifuss [Artemisia], 500-800' hoch." (cf. K. Koch, 1849: 670); "Territorium Caucasi: f. 4 (... Tatuni, Tiflis [Tbilisi])" (cf. Beck, 1890: 145, sub O. cernua f. cumana); "bei Tiflis [Tbilisi], Tatuni δ" (cf. Beck, 1930: 124, sub O. cernua var. cumana); Georgia, May 2001 by G. Schneeweiss, sub var. cumana on its host Artemisia meyeriana [Asteraceae]. SIUC, accessed 4 Mar. 2006).

Greece / Grecia

Crete / Creta: "Cr" (cf. Greuter & al., 1989: 258, sub O. cernua subsp. cumana); "Kriti with Karpathos, Kasos & Gavdhos" (cf. Domina & Raab-Straube, 2010, sub O. cumana).

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Hungary / Ungria: Hungary (cf. Barina & al., 2005).

India: "Central Tobacco Research Institute, Rajahmundry, Andhra Pradesh, India, C.A. Raju s.n. (LE)" (cf. Schlauer & al., 2003, loc. class. of O. cernua subsp. rajahmundrica); "One of the worst phanerogramous parasites of crop plants known. Occurs on tobacco in many areas in India. Known by different names in different parts of India such as 'malle1 in Andhra Pradesh and 'tokra' in North India" (photo by Integrated Pest and Diseases Management Expert Advisory System, accessed 24 Dec. 2010).

Iran / Irán: "bei Urumia [Urmia / Orumieh, West Azerbaijan Province] auf schotterigen Hügeln auf Sonchus und bei Dise-samek, im Juli 1884 (J.A. Knapp δ)" (cf. Beck, 1930: 124, sub O. cernua var. cumana); "Persia: W: Azerb.: Khvoy [Khoy / Khoi / Khuy, West Azerbaijan Province], Assadi in Rech. 44380. E: Khor.: 30 km W. Bojnurd [North Khorasan Province] versus Shahabad, 800 m, Rech. 55552" (cf. Gilli, 1976: 297, sub O. cernua var. cumana f. holosepala); "Persia: W: Ham.: Aq Bolagh-e Aqdaq [Hamadan / Hamedān Province], 2000 m, Rioux & Golv. AB-392" (cf. Gilli, 1976: 297, sub O. cernua var. cumana f. tenuisepala); Iran (cf. Tzvelev, 1981: 451, sub O. cernua subsp. cumana).

Iraq: "Syria: in deserto Singarae [Singar]..." (f. 4 leg. Haussknecht!)" (cf. Beck, 1890: 145, sub O. cernua f. cumana); "Wüste Singara [Singar]" (cf. Beck, 1930: 124, sub O. cernua var. cumana).

Israel and Palestinian territories [Palestine / Palestina]:

Israel: Israel (cf. Joel, 1988; Ish-Shalom-Gordon & al., 1993; Eizenberg & al., 2004); "Bet-Dagan Experimental Station (coastal plain) ... Kibbutz Hasolelim (lower Galilee)" (cf. Ish-Shalom-Gordon & al., 1994); "with host (Helianthus annuus), Bet She'an, Haré Gilboa (Is), 22-5-1984" (Kreutz, 1995: 91!); photo, "Kibbutz Bet Hashita Valley of Jesrael" by R. Jacobsohn, Agricultural Research Organization, accessed, 29 Jan. 2011).

Palestine / Palestina: "Hosts: ... irrigated vegetable crops in Lower Jordan Valley and Arava Valley [Arabah / HaArava / Wādī'Araba / Aravah, Great Rift Valley]" (cf. Feinbrun-Dothan, 1978: 213, sub O. cernua).

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Italy / Italia [unconfirmed]: "southern Italy" (cf. Kreutz, 1995: 88).

Jordan / Jordania:

"Hosts: ... irrigated vegetable crops in Lower Jordan Valley and Arava Valley [Arabah / HaArava / Wādī'Araba / Aravah, Great Rift Valley]" (cf. Feinbrun-Dothan, 1978: 213, sub O. cernua); Al-Khateeb & al., 2003.

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Kazakhstan / Kazakstan / Kazajistán: "Hab. in deserto soongoro - kirghisico [Soongoro-Kirghisici (Dzungaria - Kashgar)] occidentali hinc inde in locis argillosis atque in locis sabulosis ad ripam rivuli Tschaganka [Tschuganka] versus radicem montis Tschingistau [Tschingistan] [Eastern Kazakhstan / Kazakstan]" (cf. C.F. Ledebour, Ic. Pl. Fl. Ross. 4: 25 t. 390. 1833, loc. class. of Orobanche bicolor C.A. Mey.; Reuter, 1847: 32, sub Orobanche cernua var. bicolor); "Sibiriae altaicae deserto soongoro - kirghisico! (Fl. Alt., Karel. et Kiril [Karel. et Kiril Enum. pl. Fl. alt. No. 675])" (cf. C.F. Ledebour, Fl. Ross. 3: 321. 1841-1853); "Native of Altaia, in the Soongarian desert, in argillaceous and sandy soil, on the banks of the river Tschunganka, towards the foor of Mount Tschingistan" (cf. Don, 1838: 631, sub O. bicolor); "Sibir. alt. [Altaic Siberia]" (cf. Steudel, 1841: 230, sub O. bicolor); "ad flum. Lepsa in Soongaria, in salsis inter Usunbulak et Gorkoipiket (Karelin et Kiriloff!") (cf. Reuter, 1847: 32, sub Orobanche cernua var. bicolor); "Siberia altaica: in deserto soongoro - kirghisico [Soongoro-Kirghisici, Eastern Kazakhstan / Kazakstan] m. Alatau! (Schrenk pl. exs.)" (cf. C.F. Ledebour, 1841-1853: 322, sub O. cumana); "Songhoria: f. 4 ad fl. Lepsa inter Usunbulak et Gorkoi picket (Karelin et Kiriloff), inter Kocksu et Karatul, prope Tarba-gatai, Alakul et Saduk et prope Dschisdy-King (leg. Schrenk)" (cf. Beck, 1890: 145, sub O. cernua f. cumana); "Shungurei: bei Alakulj (Bienert)" [Songaria / Soogaria / Zhungaria / Dzungaria, Kazakhstan / Kazakstan]" (cf. Beck, 1930: 128, loc. class. of O. cernua var. cumana f. holosepala); "Turkestan / Türkistan" (cf. Beck, 1930: 125, sub O. cernua var. cumana); Kazakhstan (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Kyrgyzstan / Kirguistán: Kyrgyzstan (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Moldova / Moldavia: "Bessarabia" (cf. Reuter, 1847: 32, sub O. cumana); "Podolia [Podilia / Podillya / Поділля / Подолье) australi a Bessero indicata"] (cf. Reichenbach, 1862: 107, sub O. cumana); Moldova (cf. Sharova, 1968); "Ju" (cf. Greuter & al., 1989: 258, sub O. cernua subsp. cumana); "in 1981, 230 flowers of the parasite were registered on one plant in Komrat [Conrat, Gagauzia] district of Moldova in some sunflower fields" (cf. Yakutkin, 2003-2009).

Mongolia: "Mongolia: f. 4: Oasis Nija 1500 m, Oasis Keria, in decliv. N. Thianschan, prope Zaidam (leg. Przewalski)" (cf. Beck, 1890: 145, sub O. cernua f. cumana; Beck, 1930: 125, sub O. cernua var. cumana; Mongolia (Tzvelev, 1981: 451, sub O. cernua subsp. cumana; Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Morocco / Marruecos: "Also in northern Africa" (cf. Kreutz, 1995: 88).

Nepal: Nepal (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Romania / Rumanía: "Rumänien: δ Lacul sarat bei Braila, im August (Prodan) und in der Dobrudscha: δ bei Cerna voda, Tschirghial, im Juli (Prodan)" (cf. Beck, 1930: 123, sub O. cernua var. cumana); Romania (cf. Vrânceanu & al. 1980; Pacureanu-Joita & al., 1998; Gagne, 1998).

Russia / Rusia: "O. arenaria M. In arena mobili ad Wolgam versus Astrachan lecta, 1803" [Province of Astrakhan / Astrakhan Oblast, Russia] (cf. Wallroth, 1825: 58, loc. class. of Orobanche cumana; Tzvelev, 1981: 451); t. 691 f. 926 ["Caucaso: Blume, ex herb. cl.Trevirani"] f. 927 ["Specimen Wallrothianum"] (cf. Reichenbach, 1829: 45, sub O. cumana); "prope Stauropolim" [Stavropol / Stauropolis] (cf. Reuter, 1847: 32, sub O. cumana); "Hab. in Rossia australi [ad Tanain (Henning), pr. Astrachan (M. a Bieb.), in deserto caspio (Tauscher ex (M. a Bieb.)] et cumanensi (M. a Bieb.) et provinciis caucasicis [pr. Stawropol (Stavropol / Stauropolis)] (C.A. Meyer), ad fl. Terek [Terek River, loc. class. of O. sabulosa Steven]! (Steven pl. exs.)]..." (cf. C.F. Ledebour, 1841-1853: 322, sub O. cumana); "In Tatarisch [Tatarstan, Russia]-Grusien [Georgia] und Schirwan [Shirvan, Azerbaijan] auf Beifuss [Artemisia], 500-800' hoch." (cf. K. Koch, 1849: 670); "... Sarepta [Sarepta (now part of Volgograd, Volgograd Oblast), Southeast of the European part of Russia] (in Xanthio strumario) Becker!" (cf. Reichenbach, 1862: 107, sub O. cumana); "Sibiria orienttali prope Ust Balei [Irkutsk Oblast]" (Trautvetter 1871: 26, loc. class. of Orobanche glaucantha); "Caucasus ad Stavropol (Stawropol / Stauropolis)] (C.A. Mey!)" (cf.  Boissier, 1879: 545, sub O. cumana); "Rossia: f. 4 ... Gub. Astrachan (! Astrachan, Bogdo; Sarepta), Gub. Saratow (! Kamyschin), Gub. Simbirsk (! Syssan), Gub. Perm! in montibus Uralensibus!, Gub. Orenburg!" (cf. Beck, 1890: 144, sub O. cernua f. cumana); "Gub. der Donschen Kosaken: am Don [Rostov Oblast] bei Butajsk auf Xanthium spinosum (Paczoski δ) und bei Taganrog [Rostov Oblast] (Schmalhausen δ) im Kreise Ustj-Medwedizkaja und Taganrog (J. Novopokrovsky n. 746!) auf Helianthus annuus; Gub. Saratow [Saratov Oblast]: bei Saratoff, Kamyschin, Zarizyn (Persidski δ) usw.; Gub. Simbirsk [Ulyanovsk Oblast]: bei Syssan (Kolakowsky δ); Gub. Perm [Perm Krai] und im Ural; Gub. Orenburg [Orenburg Oblast]; im Gub. Astrachan [Astrakhan Oblast]: bei Astrachan, Bogdo, Sarepta [now part of Volgograd, Volgograd Oblast], Dolban an der Mündung der Wolga [Astrakhan Oblast] auf Artemisia maritima, im Juni (Paczoski δ); in Ciskaukasien: im Gub. Kuban: auf der Halbinsel Taman (Lipsky δ, γ), bei Noworossisk an sandigen Stellen im Juli (Radde δ), bei Anapa (Lipsky δ); im Distrikte Stawropol [Stavropol Krai] bei Stawropol (Lipsky δ), Sergjewskoje (Akinfiew δ) ... Daghestan [Republic of Dagestan]: bei Tezir jurt im Mai (Lipsky δ, £); im Distr. Dargi bei Ametherk, bei 1170 m Seehöhe, im Juli (Alexeenko δ)" (cf. Beck, 1930: 123-124, sub O. cernua var. cumana); «Sands in the lower reaches of the Volga and on the Kuma rivers ("In arena mobili deserti astracanici et cumani"). ... "O. arenaria M. In arena mobili ad Wolgam versus Astrachan lecta, 1803"» [Astrakhan Oblast] (cf. Tzvelev, 1981: 451, sub O. cernua subsp. cumana); Russia (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Serbia: Serbia (cf. Škorić & al. 2010: 64).

Slovakia / Eslovaquia: Slovakia (cf. Cagáň, 2006); "Slovakia" (cf. Domina & Raab-Straube, 2010, sub O. cumana).

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Spain / España (cf. Díaz Celayeta, 1974; Velayos, 1978; González Torres & al., 1982; Castejón Muñoz, Romero Muñoz & García Torres, 1989; Cubero, 1994: 466, "The presence of O. cernua Loefl. (syn. O. cumana Wallr.) in Spain is a rather recent phenomenon, the first reference dating back to 1958; Gagne, 1998; Pujadas & Velasco, 2000; Pujadas & al., 2007; Fernández Escobar & al., 2009).

Andalusia / Andalucía (López Granados & García Torres, 1992; García Torres & al, 1994: 625, f. 2).

Cádiz: Jerez ... Villamartín ... (cf. García Torres & al, 1994: 625, f. 2); "Puerto Serrano, on cultivated sunflower (Helianthus annuus L.), 3.vi.1989, J. Oliveira (COA 13524). Jerez de la Frontera, vii.1995, J. Fernández Escobar, CA195 (COA 28285)" (cf. Pujadas & Velasco, 2000: 517).

Córdoba: Bujalance ... Sta. Cruz ... (cf. García Torres & al, 1994: 625, f. 2); "Valle del Guadalquivir: de Espejo a Montemayor, c. Ermita Dos Hermanas, 30SUG5768, A. Pujadas & J. Fuertes, COA 33962" (cf. Pujadas & al., 2007: 14); "Córdoba, Finca Ruano, on cultivated sunflower, 9.vi.1993, F. Cartujo (COA 17472). Córdoba, Parque Cruz Conde, 16.vi.1994, P. Blanco (COA 17471). Córdoba, near Airport, on cultivated sunflower, 30.vi.1996, A. Lora (COA 22078). Córdoba, Huerta de San Luis, on cultivated sunflower, 30SVG4092, 2.vii.1996, A. Lora & A. Pujadas (COA 22077). Córdoba, Cortijo del Pardito, on cultivated sunflower, 30SUG4377, 16.vii.1996, A. Moyano & A. Pujadas (COA 22115). Puente Genil, vii.1994, J. Alvarado, CO194 (COA 28286). Córdoba, vii.1996, J.M. Melero, CO196 (COA 28287)." (cf. Pujadas & Velasco, 2000: 517).

Granada: "1986-89: ...O. cernua heavily infested ... sunflower area ... and Huescar (Granada province)" (cf. García Torres & al, 1994: 625); "Atarfe, on cultivated sunflower (oilseed), 14.ix.1995, A. Pujadas (COA 17629)" (cf. Pujadas & Velasco, 2000: 517).

Huelva: "Campiña de Huelva y Condado-Aljarafe: Escacena, 23-VI-2005, E. Sánchez-Gullón, COA 33961" (cf. Pujadas & al., 2007: 14).

Málaga: "1978-82: O. cernua spread ... sunflower area ... and Fuente Piedra (Malaga province; Gonzalez Torres & al., 1982)" (cf. García Torres & al, 1994: 624); Campillos ... (cf. García Torres & al, 1994: 625, f. 2); "Fuente de Piedra, [Los] Llanos de Antequera (E), 26-4-1994" (Kreutz, 1995: 90-91!); "Antequera, on cultivated sunflower (oilseed), 3.vi.1989, J. Oiveira (COA 13569). Campillos, on cultivated sunflower (confectionery), 13.vi.1989, J. Oliveira (COA 13525). Between Campillos and Antequera, 20 km, on cultivated sunflower (oilseed), 13.vi.1989, J. Oliveira (COA 13568). Near Laguna de Fuente de Piedra, on cultivated sunflower, 8.vi.1995, A. Lora & A. Pujadas (COA 17628)." (cf. Pujadas & Velasco, 2000: 517-518).

Sevilla: "1986-89: ...O. cernua heavily infested ... sunflower area ... at El Coronil (Seville province)" (cf. García Torres & al, 1994: 625); Lebrija ... Utrera ... Marchena ... Écija ... (cf. García Torres & al, 1994: 625, f. 2); "El Coronil, on cultivated sunflower, 16.vi.1986, L. García Torres (COA 13636). Morón to El Coronil, 9 km, on cultivated sunflower (oilseed), 2.vii.1987, J. Oliveira (COA 13624). El Coronil to Morón, 7 km, on cultivated sunflower (oilseed), 2.vii.1987, J. Oliveira (COA 13633). El Coronil to Utrera, 15 km, on cultivated sunflower (confectionery), 2.vii.1987, J. Oliveira (COA 13645). Osuna, 1.vii.1988, J. Oliveira, COA 28295. El Coronil to Montellano, 10 km, on cultivated sunflower (oilseed), 1.vii.1988, J. Oliveira (COA 13533). El Coronil to El Arahal, 12 km, on cultivated sunflower (confectionery), 1.vii.1988, J. Oliveira (COA 13629). El Coronil to Acabalas, 10 km, on cultivated sunflower (oilseed), 1.vii.1988, J. Oliveira (COA 13643). Montellano, on cultivated sunflower (confectionery), 13.vi.1989, J. Oliveira (COA 13618). Osuna, vii.1989, J. Oliveira (COA 28296). Écija, Sotillo Gallego, vii.1993, J.M. Melero, SE193 (COA 28290). Écija, Finca Pavía, vii.1994, J. Fernández, SE194 (COA 28293). Écija to Herrera, 24 km, on cultivated sunflower, 18.v.1995, A. Pujadas (COA 17460). Écija, Finca Mencáa, vii.1996, J.M. Melero, SE296 (COA 28292). Écija to Estepa, 3 km, 1.vii.1996, A. Pujadas, A. Lora & A. Jiménez (COA 22079). Herrera to Écija, 3 km, 1.vii.1996, A. Pujadas, A. Lora & A. Jiménez (COA 22080). Aznalcázar, near Guadiamar river, on cultivated sunflower, 29SQB4232, 22.xii.1998, A. Pujadas & M. del Río (COA 27749). Cañada del Rosal, vii.1998, J.A. Sainz de Tejada (COA 28291). Écija, viii.1998, J.A. Sainz de Tejada (COA 28294)" (cf. Pujadas & Velasco, 2000: 519-519).

Castile and León / Castilla y León (cf. Fernández Escobar & al., 2009).

Valladolid: "Medina del Campo" (cf. Fernández Escobar & al., 2009).

Castile-La Mancha / Castilla-La Mancha (cf. Díaz Celayeta, 1974; Sobrino Vesperinas & Martínez Abad, 1980; García Torres & al, 1994)

Cuenca: Cuenca (cf. Sobrino Vesperinas & Martínez Abad, 1980); "1978-82: O. cernua spread ... sunflower area of Cuenca ..." (cf. García Torres & al, 1994: 624); "Villarejo de Penesteban, viii.1992, J.M. Melero, CU192 (COA 28288). Alcázar del Rey, ix.1994, J.M. Melero, CU494, (COA 28289). Km. 17,800 of Cuenca road, 2.viii.1978, M. Velayos (MACB 47083)" (cf. Pujadas & Velasco, 2000: 517); Cuenca, Barajas de Melo, M. Velayos (cf. Flora iberica, sub O. cernua [on Helianthus annuus L.], accessed, 21 Dec. 2010); Cuenca (cf. Fernández Escobar & al., 2009); Municipality of Almendros, between la Noria and Saleguillas (at km 5, CM-3011, Almendros-Casas Luján), parasitic on Scorzonera laciniata L. in an ancient terrace-cultivation, G. Gómez Casares & G. Moreno Moral MM0047/2007, 12 May 2007 (herb. Sánchez Pedraja 12840) [N.B. this is the first record of O. cumana parasitizing a wild plant in the Iberian Peninsula].

Toledo: (cf. Díaz Celayeta, 1974; Gonzalez Torres & al., 1982; García Torres & al, 1994: 624, "1958: O. cernua was first observed in Toledo ... (Díaz Celayeta, 1974)"; Fernández Escobar & al., 2009).

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Syria / Siria: "Syria: ... et fl. Chabur [Khabur River] (f. 4 leg. Haussknecht!)" (cf. Beck, 1890: 145, sub O. cernua f. cumana); "in der Wüste Singara und am Flusse Chabur (Haussknecht δ)." (cf. Beck, 1930: 124, sub O. cernua var. cumana); "Mesopotamien: bei Tell Halaf in der Steppe bei Haus, im Mai (Kohkl! in der Expedition Frh. v. Oppenheim) [Tell Halaf, Al Hasakah]" (cf. Beck, 1930: 128, loc. class. of O. cernua var. cumana f. valens).

Tajikistan / Tayikistán: "Turkestania: prope Suidan et Pamil [Pamir] (f. 4 leg. Kuschalkewicz) ... pr. Kuldja (4 leg. Schrenk)" (cf. Beck, 1890: 145, sub O. cernua f. cumana); "in planitie Kickdan ad montes Gasi Machik et prope Gomut (f. 5 leg. Regel)" (cf. Beck, 1890: 145, loc. class. of Orobanche cernua f. tenuisepala); "Turkestan: bei Suidan umd Pamil [Pamir] (Kuschalkewicz δ!), auf der Ebene Kickdan am Gasi Machik Gebirge und bei Gomut (Regel f. 6!), bei Kuldja (Schrenk δ!)" (cf. Beck, 1930: 125, sub O. cernua var. cumana); Tajikistan (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Tunisia / Túnez: "Also in northern Africa" (cf. Kreutz, 1995: 88).

Turkmenistan / Turkmenistán / Turkmenia / Туркмения: "Siberia uralensi: in Turkomania! (Eichw., Karelin pl. exs.)..." (cf. C.F. Ledebour, 1841-1853: 322, sub O. cumana); "Turkomania (Eichw.)" (cf.  Boissier, 1879: 545, sub O. cumana); "Transcaspia: prope Krasnowok f. 4" (cf. Beck, 1890: 145, sub O. cernua f. cumana); "Transkaspien: bei Krasnowodsk" (cf. Beck, 1930: 125, sub O. cernua var. cumana); Turkmenistan (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Turkey / Turquía: "Phrygien [Phrygia]" (cf. Beck, 1930: 124); "In 1983/85 more than 50% of the Thracia [Eastern Thrace] sunflowers in northern Turkey was infested" (cf. Petzoldt & Sneyd, 1986); Turkey (cf. Bulbul & al., 1991; Gagne, 1998; Kaya & al., 2004); SW Asia (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

Ukraine / Ucrania: "In arenosis Podoliae" (cf. Reuter, 1847: 32, sub O. cernua var. bicolor); "Podolia (Podilia / Podillya / Поділля / Подолье) australi a Bessero indicata" (cf. Reichenbach, 1862: 107, sub O. cumana); "Russische Länder: Ukraine: in Bessarabien bei Tatar-Bunar (Zelenetzky δ) um Reni (Herb. Besser δ), Pandaclia (Grintescu δ); Gub. Gherson: bei Odessa (Schmalhausen, Engler (δ), Cherson (Paczoski δ), Gub. Jekaterinoslaw [Dnipropetrovsk]: um Jekaterinoslaw, im Sept. (Alexeenko δ), im Distrikte Alexandrowsk [Zaporizhia Oblast] bei Nowohupolowka, im August-November auf Helianthus und Lycopersicum (Alexeenko δ); im Distrikte Nowomoskowa, im Juli (Alexeenko δ)" (cf. Beck, 1930: 123-124, sub O. cernua var. cumana); "Ukraine" (cf. Domina & Raab-Straube, 2010, sub O. cumana).

Crimea / Krym: "Tauria [Crimea / Krym]" (cf. Reuter, 1847: 32, sub O. cumana); "Rossia mer. (Tauria)" (cf. Nyman, 1854-1855); "Sudak [Sudaq / Судак] Steven! ..." (cf. Reichenbach, 1862: 107, sub O. cumana); "Tauria (M.B.)" (cf.  Boissier, 1879: 545, sub O. cumana); "Rossia: f. 4 in Gubernia Tauria (! desertum Nogaiorum, Jalta etc.)" (cf. Beck, 1890: 144, sub O. cernua f. cumana); "in der Krim und Taurien: bei Aleschki (Paczoski ß, δ) ... Jalta, Nikita und Kertsch, im April bis Juni (Paczoski δ), um Kopsel (A. Callier δ), um Simeiis (Golde δ! auch f. 4), bei Balaklava, Inkermann, Sudak (Fedtschenko δ!) auf der Halbinsel Kercz (Klopotow δ) im Mai-Juli" (cf. Beck, 1930: 123-124, sub O. cernua var. cumana); "In der Krim [Crimea / Krym] bei Oreanda (V. Golde! in Herb. Petersburg. [LE] unter O. alba)" (cf. Beck, 1930: 128, sub O. cernua var. cumana f. taurica); "RK" (cf. Greuter & al., 1989: 258, sub O. cernua subsp. cumana); "near Yevpatoriya, Crimea [Crimea / Krym], UA, (KW)" (cf. Tzvelev, 1981: 451; view IPNI entry; Schlauer & al., 2003, loc. class. of O. sarmatica); "Crimea" (cf. Domina & Raab-Straube, 2010, sub O. cumana).

Uzbekistan / Uzbekistán: "Chiwa [Xiva / Хива, Choresmien / Xorazm / Choresm Province]: (4 leg. Korolkow et Krause)" (cf. Beck, 1890: 145, sub O. cernua f. cumana; Beck, 1930: 125, sub O. cernua var. cumana); "Bokhara [Bukhara]: in monte Taschbulak district. Hissar (f. 4 leg. Regel)" (cf. Beck, 1890: 145, sub O. cernua f. cumana); Uzbekistan (cf. Zhang & Tzvelev, 1998, sub O. cernua var. cumana).

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