Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

ISSN: 2386-9666 - Last Updated: 2 May 2018


Accepted Name:

Diphelypaea tournefortii (Desf.) Nicolson in Taxon, 24(5-6): 654 (1975 [Nov 1975])

Basionymum / Basionym:

Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298[-300], t. 21 (1807) (Desfontaines, 1807: 298-300, t. 21; 1808, 16, t. 10; Sprengel, 1825: 818; Don, 1838: 633)

Synonyms:

Homotypic synonyms:

Phelypaea Orientalis, flore coccineo Tournef., Corollarium: 47 t. 479 (1703) [pre-Linnaean name, not validly published] (Desfontaines, 1807, 298 "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Desfontaines, 1808, 16 "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Don, 1838: 633; Stapf, 1915: 285; Cullen, 2010: 380)

Lathraea phelypaea var. β [unnamed] L., Sp. Pl. 2: 606 (1753 [1 May 1753]) ["β Orobanche orientalis, flore coccineo. Tournef. cor. 47."], nom. illeg. (Stapf, 1915: 286)

Anoplon tournefortii (Desf.) G. Don, Gen. Hist. 4: 633 (1837-1838)

Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) ["A. Tournefortii Walp. rep. bot. 3 p. 48, ex parte"]; [Walp., Repert. Bot. Syst. 3: 481 (1844 [12-14 Sep 1844]) (Beck, 1930: 44)]. Obs. "basi vaginis elongatis inaequalibus involuti. Calyx ... villosus ... (v. s. in h. Bentham.)" (Reuter, 1847: 43)

Aeginetia tournefortii (Desf.) [Juss. sec. Steudel] Beck in Engl., Pflanzenr. 96: 44 (1930), pro syn.

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Heterotypic synonyms:

? = Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name, not validly published]. Syn.:  Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]) (Linnaeus, 1753: 606; Don, 1838: 635); = Phelypaea foliata Lamb. in Trans. Linn. Soc. London 10: 260 t. 7 (1811) (Endlicher, 1839: 727)

? = Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]). Syn.:  Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name] (Linnaeus, 1753: 606; Don, 1838: 635);  Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838); = Phelypaea foliata Lamb. in Trans. Linn. Soc. London 10: 260 t. 7 (1811). Ind. loc.: "Habitat in Oriente" (Linnaeus, 1753: 606); "Asia minor" (Sprengel, 1825: 818); "H. Native of the Levant" (Don, 1838: 635; Lathraea anblatum in the Linnaean Typification Project, accessed Mar 2012). Obs. According to Endlicher (1839: 727), Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) is conspecific with Phelypaea foliata Lamb. in Trans. Linn. Soc. London 10: 260 t. 7 (1811). According to Reuter (1847: 43), Anblatum tournefortii G. Don is conspecific with Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807). According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753) [this synonym seems correct on the base of Don's description]

? = Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838) (Reuter, 1847: 43). Obs. According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753)

? = Aeginetia coccinea Juss. ex Reut. in DC., Prodr. 11: 44 (1847), pro syn. Syn.: = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 44 (1847) (Reuter, 1847: 44)

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= Anoplanthus coccineus var. peduncularis Boiss., Fl. Orient. 4(2): 494 (1879 [Apr-May 1879]) ["A. coccineus β peduncularis"]. Syn.: = Phelypaea orientalis, flore coccinea Tournef., Corollarium: 47 t. 479 (1703); = Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807); = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) (Boissier, 1879: 494; Beck, 1930: 44; Novopokrovskii & Tzvelev, 1958[2000]: 28[35]). Ind. loc.: "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, Turkey] (Tourn!, Huet!, Calv!), in Kurdistaniâ (Barré de Lancy [vice-consul de France à Mersine et Adana]!, Armeniâ Rossicá (Buhse!), in pratis montosis Georgiae Caucasicae (Hoh!)". Ic.: "Phelypaea Tournefortii Desf. Cor. tab. 10" (Boissier, 1879: 494). Obs. "Squamae ad basin caulis superne longe nudis magis congestae" (Boissier, 1879: 494)

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Ind. loc.: "Il est originaire d'Arménie [Western Armenia]" (Desfontaines, 1807: 300); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum [Erzurum, Anatolia, Turkey] (Tournefort..." (Beck, 1930: 44; Novopokrovskii & Tzvelev, 1958[2000]: 28[35]; Nicolson, 1975: 652 "variety B [β]"; Cullen, 2010: 380; view IPNI entry).

Typus / Type: "Type: Describibed from Turkey (P)" (Novopokrovskii & Tzvelev, 1958[2000]: 28[35]); "Type: Phelypaea orientalis, flore coccinea, Tournefort [t. 479]" (Gilli, 1982: 2); "that no specimens of the Phelipaea Tournefortii now remain in his Herbarium, or are known to exist in any other collection" (Lambert, 1811: 261); "No specimen of this species could be found in the Tournefort herbarium (P)" (Gilli, 1982: 2); "There is, apparently, no specimen in the Tournefort herbarium, but Aubriet, the artist who accompanied Tournefort, made an illustration which appears as t. 479 of the Corollarium, and can be regarded as the type material; it is clearly the plant recognised here as Phelypaea tournefortii." (Cullen, 2010: 380).

Designated by: D.H. Nicolson in Taxon 24: 652 (1975 "variety B [β]").

Host: Parasitic exclusively on Tanacetum species (Asteraceae / Compositae) (Annotated Checklist of Host Plants of Orobanchaceae, accessed Aug 2009); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. [?]" (Reuter, 1847: 43; Beck, 1930: 44; Novopokrovskii & Tzvelev (1958[2000]: 28[35]); "On Achillea and Tanacetum" (Gilli, 1982: 2).

Comments:

Morphological observations: "P. Scapo basi involuto, villoso ... Calice ... velues ... Le Phelipaea de Tournefort a les tiges nues et les divisiones de la corolle sont circulaires et non ovales" (Desfontaines, 1807: 298-300). Key to these species (Stapf, 1915: 290; Novopokrovskii & Tzvelev (1958[2000]: 26[33]; Cullen, 2010: 383-384); pollen morphology (Zare & al. 2014).

Phylogenetic observations: Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57 [x = 19, sub Phelypaea (syn. Diphelypaea)], 61 f. 4 ["Phylogenetic relationships", sub Phelypaea], 63 (2005); "Recent molecular phylogenetic studies suggest two major lineages, informally called the Orobanche-group and the Phelipanche-group (Schneeweiss et al., 2004a). The first includes Orobanche sect. Orobanche and the small SW Asian genus Phelypaea (listed as Diphelypaea in Schneeweiss et al., 2004a), while the second includes the remaining sections of Orobanche (sects. Gymnocaulis, Myzorrhiza, and Trionychon). The circumscription of these groups is well supported by the distribution of chromosome base numbers, these being x = 19 in the Orobanche-group and x = 12 in the Phelipanche-group (Schneeweiss et al., 2004b)" Weiss-Schneeweiss & al., 2006; Schneider, 2016: f. 1.

Taxonomic observations: Diphelypaea tournefortii (Desf.) Nicolson in Taxon, 24(5-6): 654 (1975), "without full basionym ref." view IPNI entry. The reference to the place of publication of the basionym in "References"[!] (Nicolson, 1975). Phelypaea tournefortii in MBG - w3TROPICOS, accessed 6 Feb 2006. Phelypaea Hill in Owen, Dict. Arts Sci. 3: 2416 (1755 [1 Mar 1755]) (Reveal, 1991: 228, "The Hill genus name is now the oldest one available")

Chromosomatic number: n = 19, 2n = 38 (Schneeweiss & al., 2004, sub Diphelypaea; Carlón & al., 2005: 57, sub Phelypaea [between Bayburt and Erzurum, NE Anatolia, Turkey]).

Ic.: a, Tournefort, Corollarium: 47 t. 479 (1703, sub Ph. orientalis, flore coccinea); b, Desfontaines 1807: t. 21 et 1808: 16, t. 10, sub Phelypaea tournefortii; c, Beck in Engl., Pflanzenr. 96: 43 f. 9 (1930, sub Ph. Tournefortii); d, habit (Muradiye, Van, Turkey [May 2009, photo by P. Coulot]); e, flower (ib.); f, calyx (ib. ["Calyx ... villosus" (Reuter, 1847: 43)]); g, scale-leaves (ib. ["Squamae ad basin caulis superne longe nudis magis congestae" (Boissier, 1879: 494)]); h, habit (Ph. tournefortii eastern Anatolia, Jul 2002 by G. Schneeweiss); i, flower (Ph. tournefortii eastern Turkey, by L.J. Musselman); j, sheet (Turkey, by Demirkuş, 2007); k, Novopokrovskii & Tzvelev (1958[2000]: 30[37] t, 3 f. 5); l, D. tournefortii vs. D. coccinea vs. D. boissieri (P, habits and calyces).

Distribution: Eurasian distribution (Armenia, Azerbaijan, Georgia, Turkey). Eurasian distribution (Domina & Raab-Straube, 2010, sub Phelypaea tournefortii, accessed Jul 2010).

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Specimens examined and literature records.

Armenia / Hayastan / Armenien / Arménie / Армения

Ararat Province / Араратская область: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / В 2 км к югу от с. Кярки [2 km to the south of the village of Karki / Kərki / Kyarki / Tigranashen / Тигранашен (Садаракский район / Sadarak District)] можжевеловое редколесье, 1400 м. / 29 мая 1973 г. [29.05.1973] Н. Шведчикова [N. Shvedchikova]" (MW 0723306; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.); "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / Окрестности Советаmена [Surroundings of the Council (Vedi)]. / Н. Шведчикова [N. Shvedchikova] 30 мая 1973 [30.05.1973] г. 7782" (MW 0723307; rev. Ó. Sánchez Pedraja Apr 2018); "Plants of Armenia / ... / Flile NY: 288.0 / Phelipaea coccinea (M. Bieb.) Poir. / det. K. Tamanyan, / Armenia. 3 km S from Lusashoh village [Lusashogh / Garaxach / Лусашох], 39º49'72N, 44º59'195E / 1760 m. (... / ...). Mountain steppe. / Kamilla Tamanyan 10-2005  28 May 2005 / George Fayvush / The New York Botanical Garden (NY) / in collaboration with / Institute of Botany, Armenian Academy of Sciences (ERE)" (P 03931901; rev. Ó. Sánchez Pedraja Apr 2018); "Цветущее растение. Армения, Гарни, Хосровский заповедник [Garni, Khosrov Reserva] 18.05.2010" (photo 107519 by Ваэ Мартиросян [Vahe Martirosyan], 2011 in Plantarium [Diphelypaea tournefortii, on Tanacetum sp.] and iNaturalist.org; accessed Feb 2013); "Khosrov reserve 2295 m a.s.l.", 5 June 2007 (photo 744571 by Roberto Sindaco, sub Diphelypaea, in iNaturalist.org; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [the photo seems to correspond to D. tournefortii on Tanacetum sp.).

Gegharkunik Province / Гехаркуникская область: "im Distr. Novobajazet [Novo Bajazet / Nor Bayezet / Novyi Bayazet, Ajridzha / Armaghan Lerr] um Solak, Berezovaja butka, Konstantinowska (E. Mylow!), bei Station Eilar [Dzak] (A. Fomin!)" (Beck, 1930: 44, sub P. tournefortii); "Diphelypaea tournefortii, filmed in the Tarachichaki [Tsaghkunk/ Varazhnunik] (town) ..." (photo 85148 by Ваэ Мартиросян [Vahe Martirosyan], 2011 in vahemart.livejournal.com; accessed Jan 2015).

Kotayk / Kotaik Province / Котайкская область: "Herbarium Universitatis Mosquеnsis / Plantae Caucasicae Collaboratoribus Horti / Botanici Universitatis Mosquеnsis Lectaе / Phelураea tournеfortii Desf. / Армения, Разданский р-н [Hrazdan raion], / хр. Цахкунец [Tsaghkunyats Mountains], басс. р. Мармарик [Marmarik], / окр. пос. Цахкадзор [Tsaghkadzor / Tsakhkadzor] / М.Г. Пименов Е.В. Клюйков / Л.П. Томкович Н.А. Кириллина [M.G. Pimenov, E.V. Klyuykov / L.P. Tomkovich, N.A. Cyrillin] / 5 июня 1977 [5 June 1977]" (MW 0723305; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.); "Diphelypaea tournefortii, Berdatak, 2010.05.18 // Tsaghkadzor [Tsakhkadzor / Darachichag]" (photos 5788747528409755042 and 179759 by Vahe Martirosyan [Ваэ Мартиросян] in Flora of Armenia's Check List and iNaturalist.org; accessed Feb 2013); "Цветок. Армения, область Котайк, окрестности Цахкадзора [neighborhood Tsakhkadzor] 31.05.2008" (photo 107214 by Ваэ Мартиросян [Vahe Martirosyan], 2011 in Plantarium [Diphelypaea tournefortii]; accessed Feb 2013); "Kotayk' [Yelgovan], Armenia" (photo 57543824 by Vahe Martirosyan in www.panoramio.com, accessed Mar 2012); "Цветущее растение. Армения, обл. Котайк, окр. села Вохчаберд, разнотравная степь [Armenia, reg. Kotayk, env. Voghjaberd village, grassland steppe]. 26.05.2012" (photos 166927 et 166926 by Владислав Куропаткин [Vladislav Kuropatkin] in Plantarium [Diphelypaea tournefortii]; accessed Feb 2013); "Diphelypaea tournefortii, Tsaghkadzor, 2014.06.04" (photo 6024440769631112354 by Vahe Martirosyan [Ваэ Мартиросян] in http://vahemart.livejournal.com; accessed Jan 2015); "Photo taken in Jrvezh Forest Park, H-3, Jrvezh, Armenia 40° 10' 35.44" N  44° 37' 51.03" E" (photo 3909676 by in www.panoramio.com; accessed Jan 2015); "Armenia, near Tseghkadrov city, dry grassland 40,524303º N ; 44722640º E. 1 5.05.2014. M. Peregrym" (Леонідович [Leonidovich], 2016: 106, sub Phelypaea tournefortii).

Vayots Dzor Province / Вайоцдзорская область: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армянская ССР, Даралaгез [Armenian SSR, Daralagez]. Гора Теке-Долдуран, урочище Xазма [T’ek’sari Lerrnashght’a range?, natural boundary Hazma?], / горные луг [mountain meadow], 10 июля 1929 г. [10.07.2019] / Собрали Ант. и И.llyкины [Leg. Ant. and I. llykin]" (MW 0723304; rev. Ó. Sánchez Pedraja Apr 2018); "Цветки. Армения, Селимский перевал [Selim Pass / Orbelian's Caravanserai]. 20.05.2014" (photo 238873 by Тамара Галстян, 2014 in Plantarium [Diphelypaea tournefortii]; accessed Jan 2015); "Цветок. Армения, Вайоц Дзор, окр. монастыря Спитакавор [Vayots Dzor, env. Spitakavor monastery]. 02.05.2013" (photos 188448 et 188447 by Марина Скотникова, 2013 in Plantarium [Diphelypaea tournefortii]; accessed Jan 2015).

Yerevan / Erywan / Jerewan / Ереван: "Caucasus, Gouv. Eriwan, St. Eyliar, on Pyrethrum myriophyllum, Thomin (in Acta Hort. Tifl. vi. II. (1902) 56)" (Stapf, 1915: 293, sub Anoplanthus coccineus var. nigrovittatus) [Pyrethrum myriophyllum is the typical host of D. tournefortii]; "Distr. Eriwan [Erywan / Yerevan / Jerewan / Ереван]: bei Hilar, zwischen Garyna und Arabat (Bayern)" (Beck, 1930: 44, sub P. tournefortii).

Unknown location: "Herbarium Universitatis Mosquеnsis (MW) / P. Smirnow. Pl. Armeniae ross. a. 1929. / Anoplanthus coccineus Walp. / ... / Daparuras / ... / 19.VI.1929 / N.º 743 ..." (MW 0723296; rev. Ó. Sánchez Pedraja Apr 2018); "Velmi výraznými prvky jsou zástupci zárazovitých (Orobanchaceae), např. nepřehlédnutelná rostlina s jediným červeným květem Phelypaea coccinea (obr. 10), která parazituje na chrpách (Centaurea)." (Ekrtová & Ekrt, 2012: 70, 72, photo 10; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [the photo seems to correspond to D. tournefortii]).

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Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaïdjan (Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); Stapf, 1915: 288, sub Phelypaea orientalis, flore coccinea; Beck, 1930: 44, sub P. tournefortii; Karjagin, 1957)

Khizi / Xızı Rayon / Хызинский район: "Anoplon Biebersteinii ... // Armenia Rossica [Dübrar Dağı?, near Altıağac / Alty-Agatsch?] / Seidlitz [N.K. von Seidlitz] / Herb. Al. de Bunge" (P 02970936 - E. Cosson, rev. Ó. Sánchez Pedraja Nov 2011 [three specimens on the lover left corner close to Bunge's labels]); "Russian Armenia" (Stapf, 1915: 288, sub Phelypaea orientalis, flore coccinea); "Calculated" (Domina & Raab-Straube, 2010, sub P. tournefortii).

Khojavend / Xocavənd Rayon / Ходжавендский район [unconfirmed]: "Tale Dschagritschai, bei Takjalta Kulu [Dağdöşü / Tak] (Bayern!)" (Beck, 1930: 44, sub P. tournefortii).

Nakhchivan Autonomous Republic / Нахичева́нская Автоно́мная Респу́блика / Naxçıvan Muxtar Respublikası

Бабекский район / Babek Rayon: "444. Anoplanthus Tournefortii / Walp. ex parte / Puskoh bei Nachitschewan / 31/V, 47 [31 May 1847] / Buhse [leg.] // Iter Persicum. / Buhse, 1847-1849 / Herb. Al. de Bunge" (P 02970936 - E. Cosson, rev. Ó. Sánchez Pedraja Nov 2011 [the two specimens on the upper left corner close to Buhse's label]); "Armeniâ Rossicá (Buhse!)" (Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Transcaucasia: Near Pusko in the Dshagri valley [Dzhagry], north of Nahitshewan [Nachitschewan / Nakhchivan / Нахичевань], on Pyrethrum myriophyllum, C.A. Mey., Buhse" (Stapf, 1915: 293, sub P. tournefortii); "in der Provinz Nachitschevan [Nakhchivan Autonomous Republic] um Nachitschevan [Nakhchivan / Naxçıvan] bei Puskoh (Buhse!)" (Beck, 1930: 44, sub P. tournefortii); "Herbarium Universitatis Mosquеnsis (MW) / ... / Filiatonis Azerbajdzhanicae Academiae Scientiarum URSS / Herbarium Azerbajdzhanicum / Phelураea coccinea Poir. / Transcaucasia, Azerbajdzhan, respublica Nachitshevan, distr. / Nachitshevan, prope pag. Buzgov [Aşağı Buzqov]. / in calcareis 29.V.1939 Leg. E. Chalivov / Det. ..." (MW 0723302; rev. Ó. Sánchez Pedraja Apr 2018).

Садаракский район / Sadarak Rayon: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / В 2 км к югу от с. Кярки [2 km to the south of the village Karki / Kərki / Kyarki / Tigranashen / Тигранашен] можжевеловое редколесье, 1400 м. / 29 мая 1973 г. [29.05.1973] Н. Шведчикова [N. Shvedchikova]" (MW 0723306; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.).

Шарурский район / Sharur Rayon: "Herbarium Universitatis Mosquеnsis / Azərbaycan Mərkəzi Herвarium /... / Phelураea Tournеfortii Desf. / Transcaucasia, respublica Naсhitschevan, prope p. / Karakush, in herbosis. / Leg L. Prilipkо / 31-V-1934 Det. L. Prilipkо" (MW 0723308; rev. Ó. Sánchez Pedraja Apr 2018).

Unknown location: "Herbarium Universitatis Mosquеnsis (MW) / ... / Phelураea coccinea Poir. / ... [Кавказ, Азербайджан / Caucasus, Azerbaijan] / 10.VI.1931" (MW 0723298; rev. Ó. Sánchez Pedraja Apr 2018).

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Georgia / Sakartvelo / Georgien / Géorgie [needs be confirmed]

"Anoplon Biebersteinii C.A.M. / Caucas / 1850. / leg. Kovalevsky" (P 02970929, rev. Ó. Sánchez Pedraja Nov 2011 [only one specimen well preserved, "calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes; scale-leaves concentrated towards the bases of the stems" (Cullen, 2010: 384)]); "Georgia" (Tarkhnishvili & Chaladze, 2013 in Georgian Biodiversity Database, accessed Dec 2010 and Oct 2017).

Note: The gathering of Hohenaker in May 1831 and 1834 seem to correspond to D. coccinea: "pratis montosis umbrosis Georgiae caucasicae (Hohenacker !)" (Reuter, 1847: 42, sub Anoplanthus biebersteinii Reut.); "in pratis montosis Georgiae Caucasicae (Hoh!)" (Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Georgien (Hohenacker nach Boissier)" (Beck, 1930: 44).

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Turkey / Türkiye / Tuquía / Turkije (Distribution of The Taxon Over Turkey in Turkish Plants Data Service (TÜBİVES) Version 2.0 BETA; accessed Sep 2017)

Eastern Anatolia Region (Doğu Anadolu Bölgesi) "Phelypaea tournefortii: IMG_3551, IMG_3555, IMG_3579 [on Tanacetum sp.] (photos by Dana Cromie in Eastern Turkey, May 2011 [https://www.flickr.com]; accessed Feb 2013)

Northeast Anatolia Region / Kuzeydoğu Bölgesi

Ağrı Province: "B9 Ağri: Murat valley, 1600 m, D. 44038!" (Gilli, 1982: 2, sub Ph. tournefortii).

Bayburt Province / Bayburt ili: "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia" (P 02970936 - E. Cosson, rev. Ó. Sánchez Pedraja Nov 2011 [only one specimen on the lover left corner close to Huet de Pavillon's label]); 1. idem. 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (P 02970941, rev. Ó. Sánchez Pedraja Nov 2011 [six specimens very well preserved]); "Bayburt" (Distribution of The Taxon Over Turkey in Turkish Plants Data Service (TÜBİVES) Version 2.0 BETA; accessed Sep 2017).

Erzurum Province [Western Armenia]: Armenia (Desfontaines 1807: t. 21; 1808: 16, t. 10 and P02970942, sub Phelypaea tournefortii); "native of Armenia" (Stapf, 1915: 285, 288, sub Phelypaea orientalis, flore coccinea); "H. Native of Armenia" (Don, 1838: 633, sub Anoplon tournefortii); "In Armeniâ (Tournef.)" (Reuter, 1847: 43, sub Anoplanthus tournefortii); "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia" (P 02970936 - E. Cosson, rev. Ó. Sánchez Pedraja Nov 2011 [only one specimen on the lover left corner close to Huet de Pavillon's label]); 1. idem. 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (P 02970941, rev. Ó. Sánchez Pedraja Nov 2011 [six specimens very well preserved]); "Anoplon Tournefortii / ou / Phelipaea coccinea // 55. Top Dagh pres [?] / Erzeroum [Erzurum], Arménie. / Simon, 1854" (P 02970940 - E. Cosson, rev. Ó. Sánchez Pedraja Nov 2011 [seven specimes well preserved]); 1. "Orobanchea / Anoplon Tournefortii / Syn.:? / Phelipaea coccinea?". 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (P 02970943, rev. Ó. Sánchez Pedraja Nov 2011); "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, terra class. of Ph. tournefortii Desf.] (Tourn!, Huet!, Calv!)" (Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "In Kurdistaniâ [Turkish Kurdistan] (J. Brant! Strangways in h. Benth.!)" (Reuter, 1847: 43, sub Anoplanthus tournefortii); "Kurdistaniâ (Barré de Lancy [vice-consul de France à Mersine et Adana])!" (Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Herbier de Ch. d'Alleizette / ... / Phelypaea coccinea / ... env. Erzurum / Juillet 1904" (L 2821536; rev. Ó. Sánchez Pedraja Oct 2017 [cf.]); "discovered in Armenia, probably in the neighbourhood of Erzerum by Tournefort in 1701 [June 15 to July 6]" (Stapf, 1915: 285, 289, 293, sub Phelypaea orientalis, flore coccinea and sub P. tournefortii); "Turkish Kurdistan" (Stapf, 1915: 288, sub Phelypaea orientalis, flore coccinea); "Turkish Armenia: Upper Chorok valley, near Masut [Çakılı], E. of Baibut [Baiburt / Bayburt], 1200-1800 m, Huet de Pavillon! Near Erzerum, Zohrab, 826! Calver and Zohrab, 158! Kurdistan; without precise locality, Brant! Barre de Lancy (Hb. Gaillardot, 2555)" (Stapf, 1915: 293, sub P. tournefortii); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum (Tournefort, Calvert nach Boissier); im Tale Kussuklu zwischen Baiburt und Erzerum bei 4000-5000' (Huet de Pavillon!) ... Kurdistan (Barre de Lancy nach Boissier)" (Beck, 1930: 44, sub Ph. tournefortii); "E. Anatolia. A8 Erzurum: Sari Koma pass, 2600 m, Furse & Synge 850! Gümüşane: Kussuklu valley, Bayburt to Erzurum, 1220-1525 m, Huet." (Gilli, 1982: 2, sub Ph. tournefortii); "AY209285: Turkey, NE Anatolia, between Bayburt and Erzurum, between the village Maden and Kop Geçidi, 1840 m, N 40° 04' 49'' E 40° 25' 34'', on Tanacetum sp. (Asteraceae), G. Schneeweiss & C.-G. Jang (WU: S&T 7764) (Carlón & al., 2005: 63 sub Ph. tournefortii Desf.)"; "Gümüşane, Mescit Daglari: an der Straße zwischen Bayburt und Erzurum, zwischen dem Ort Maden und dem (Pass) Kop Geçidi [Bayburt Province]; 40°04'49'' N, 40°25'34'' E. 1840 m, Schneeweiß, G. & Jang, C.-G. 7764 - AY209285, 2002-06-15" (WU 32431); "Erzurum" (photo 69109 by Hayati Keser in www.treknature.com, accessed Mar 2012); "Cat. 2, 1e prijs, Rinus Bode, Phelypaea tournefortii, Kaskargebergte [Pontisch Gebergte / Kaçkar Dağları / Pontic Mountains / Kuzey Anadolu Dağları / North Anatolian Mountains], N-O Turkije" (photo 01 by NRV - Nederlandse Rotsplanten Vereniging - fotowedstrijd van 2012; accessed Feb 2013); "Fotografováno dne 8. 6. 2013 (Turecko, oblast Aşkale [Erzurum Province], na úbočí Kop Geçidi [Bayburt Province])" (photos, sub P. tournefortii, Autor Eva Rencová, 30.6.2013 in Botany.cz; 2007-2014; accessed Jan 2015).

Kars Province: "im Distr. Kaghyzman [Kagizman / Kağızman (Armenian Highlands)] am Sattel Kara dagh (S. Michailowsky)" (Beck, 1930: 44, sub Ph. tournefortii); "A9 Kars: Yağmurlu Da., 2200 m, D. 30822!" (Gilli, 1982: 2, sub P. tournefortii).

Central East Anatolia Region / Ortadoğu Anadolu Bölgesi

Malatya Province / Malatya ili: "Malatya dolaylarında çekilmiştir [it was taken near Malatya]" (photos 1/3-2/3-3/3 (53) sub Phelypaea coccinea in Türkiye'nin Çiçekleri - Yabani Çiçekler; accessed et rev. Ó. Sánchez Pedraja Oct 2017 [the three photos seem to correspond to D. tournefortii ("Stems and leaves covered with dense to sparse, long, multicellular white hairs; calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes" (Cullen, 2010: 384)), moreover we can observe the host-plant [Tanacetum sp.] in the photo 2/3]).

Tunceli Province / Tunceli ili [formerly Dersim Province]: "Tunceli, S¸Y 3245" (Zare & al. 2014, sub D. tournefortii).

Van Province: "zwischen Van und Hokotz-vank (D. Tchitouny nach Aznavour 1917)" (Beck, 1930: 44, sub P. tournefortii); "Van: between Van and Hokotz church, Tchitouny" (Gilli, 1982: 2, sub P. tournefortii); "June 13, 2002. Near Caldiran [Van], eastern Turkey" (sub Ph. coccinea, 2006 Old Dominion University, accessed 11 Dec 2010); Van: steppes le long du chemin d’accès aux chutes de Muradiye, 1826 m, UTM 38S 0392562 / 4324169, leg. P. Coulot & al., 24.05.2009 [habit, flower, calyx, scale-leaves]; Lake Van / Phelypaea tournefortii - parasite on Tanacetum species (photos 0012090 in http://www.fands.org.uk; accessed Feb 2013).

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