Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

Accepted Name:

Diphelypaea tournefortii (Desf.) Nicolson in Taxon, 24(5-6): 654 (1975)

Basionymum / Basionym:

≡ Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807) (cf. Desfontaines, 1807, 10: 298, t. 21; 1808, 16, t. 10; Don, 1838: 633)

Synonyms:

≡ Phelypaea orientalis, flore coccinea Tournef., Corollarium: 47 t. 479 (1703), nom. illeg. (cf. Desfontaines, 1808, 16 "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Don, 1838: 633; Stapf, 1915: 285)

≡ Lathraea phelypaea var. ß [unnamed] L., Sp. Pl.: 606 (1753) ["ß Orobanche orientalis, flore coccineo. Tournef. cor. 47."], nom. illeg. (cf. Stapf, 1915: 286)

≡ Anoplon tournefortii (Desf.) G. Don, Gen. Hist. 4: 633 (1837-1838)

≡ Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) ["A. Tournefortii Walp. rep. bot. 3 p. 48, ex parte"]; [Walp., Repert. Bot. Syst. 3: 481 (1844 [12-14 Sept. 1844]) (sec. Beck, 1930: 44)]. Obs. "basi vaginis elongatis inaequalibus involuti. Calyx ... villosus ... (v. s. in h. Bentham.)" (cf. Reuter, 1847: 43)

≡ Aeginetia tournefortii (Desf.) [Juss. sec. Steudel] Beck in Engl., Pflanzenr. 96: 44 (1930), pro syn.

= Anoplanthus coccineus var. peduncularis Boiss., Fl. Orient. 4(2): 494 (1879 [Apr.-May 1879]) ["A. coccineus ß peduncularis"]. Syn.: = Phelypaea orientalis, flore coccinea Tournef., Corollarium: 47 t. 479 (1703); = Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807); = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) (cf. Boissier, 1879: 494; Beck, 1930: 44; Novopokrovskii & Tzvelev, 1958[2000]: 28[35]). Ind. loc.: "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, Turkey] (Tourn!, Huet!, Calv!), in Kurdistaniâ (Barré de Lancy [vice-consul de France à Mersine et Adana]!, Armeniâ Rossicá (Buhse!), in pratis montosis Georgiae Caucasicae (Hoh!)". Ic.: "Phelypaea Tournefortii Desf. Cor. tab. 10" (cf. Boissier, 1879: 494). Obs. "Squamae ad basin caulis superne longe nudis magis congestae" (cf. Boissier, 1879: 494)

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Ind. loc.: "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum [Erzurum, Anatolia, Turkey] (Tournefort..." (cf. Beck, 1930: 44; Novopokrovskii & Tzvelev, 1958[2000]: 28[35]; D.H. Nicolson, 1975: 652 "variety B [ß]"; view IPNI entry).

Typus / Type: "Type: Describibed from Turkey (P)" (cf. Novopokrovskii & Tzvelev, 1958[2000]: 28[35]); "Type: Phelypaea orientalis, flore coccinea, Tournefort [t. 479]" (sec. Gilli, 1982: 2); "that no specimens of the Phelipaea Tournefortii now remain in his Herbarium, or are known to exist in any other collection" (cf. Lambert, 1811: 261); "No specimen of this species could be found in the Tournefort herbarium (P)" (cf. Gilli, 1982: 2).

Designated by: D.H. Nicolson in Taxon 24: 652 (1975 "variety B [ß]").

Host: Parasitic exclusively on Tanacetum species (Compositae); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec." (cf. Reuter, 1847: 43; Beck, 1930: 44; Novopokrovskii & Tzvelev (1958[2000]: 28[35]); "On Achillea and Tanacetum" (sec. Gilli, 1982: 2) (cf. Annotated Checklist of Host Plants of Orobanchaceae, accessed 24 August 2009).

Comments: "without full basionym ref." view IPNI entry (searchpage IK). The reference to the place of publication of the basionym in "References"! (cf. Nicolson, 1975).

Key to these species (Stapf, 1915: 290; Novopokrovskii & Tzvelev (1958[2000]: 26[33]; Cullen, 2010: 383-384).

Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57 [x = 19, sub Phelypaea (syn. Diphelypaea)], 61 f. 4 ["Phylogenetic relationships", sub Phelypaea], 63 (2005).

SIUC (Parasitic Plant Connection), accessed 6 Feb. 2006.

Phelypaea tournefortii in MBG - w3TROPICOS, accessed 6 Feb. 2006.

Chromosomatic number: n = 19, 2n = 38 (cf. Schneeweiss & al., 2004, sub Diphelypaea; Carlón & al., 2005: 57, sub Phelypaea [between Bayburt and Erzurum, NE Anatolia, Turkey]).

Ic.: a, Tournefort, Corollarium: 47 t. 479 (1703, sub Ph. orientalis, flore coccinea); b, Desfontaines (1808: 16, t. 10, sub Phelypaea tournefortii); c, Beck in Engl., Pflanzenr. 96: 43 f. 9 (1930, sub Ph. Tournefortii); d, habit (Muradiye, Van, Turkey [May 2009, photo by P. Coulot]); e, flower (ib.); f, calyx (ib. ["Calyx ... villosus" (cf. Reuter, 1847: 43)]); g, scale-leaves (ib. ["Squamae ad basin caulis superne longe nudis magis congestae" (cf. Boissier, 1879: 494)]); h, habit (Ph. tournefortii eastern Anatolia, July 2002 by G. Schneeweiss); i, flower (Ph. tournefortii eastern Turkey, by L.J. Musselman); j, sheet (Turkey, by Demirkuş, 2007); k, Novopokrovskii & Tzvelev (1958[2000]: 31[37] t. 3 f. 5); l, D. tournefortii vs. D. coccinea vs. D. boissieri (P, habits and calyces).

Distribution: Eurasian distribution (Armenia, Azerbaijan, Georgia, Turkey); Eurasian distribution (sec. Domina & Raab-Straube, 2010, sub Phelypaea tournefortii, accessed 20 July 2010).

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Specimens examined and literature records.

Armenia / Hayastan

Gegharkunik: "im Distr. Novobajazet [Novo Bajazet / Nor Bayezet / Novyi Bayazet, Ajridzha / Armaghan Lerr] um Solak, Berezovaja butka, Konstantinowska (E. Mylow!), bei Station Eilar [Dzak] (A. Fomin!)" (cf. Beck, 1930: 44, sub Ph. tournefortii).

Kotayk / Kotaik: "Kotayk' [Yelgovan], Armenia" (photo by Vahe Martirosyan in www.panoramio.com, accessed, 4 March 2012).

Yerevan / Erywan / Jerewan: "Caucasus, Gouv. Eriwan, St. Eyliar, on Pyrethrum myriophyllum, Thomin (in Acta Hort. Tifl. vi. II. (1902) 56)" (cf. Stapf, 1915: 293, sub Anoplanthus coccineus var. nigrovittatus) [Pyrethrum myriophyllum is the typical host of D. tournefortii]; "Distr. Eriwan [Erywan / Yerevan / Jerewan]: bei Hilar, zwischen Garyna und Arabat (Bayern)" (cf. Beck, 1930: 44, sub Ph. tournefortii).

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Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán

"Anoplon Biebersteinii ... // Armenia Rossica / Seidlitz [N.K. von Seidlitz] / Herb. Al. de Bunge" (P 02970936 - E. Cosson, rev. Sánchez Pedraja, 30 Nov. 2011 [three specimens on the lover left corner close to Bunge's labels]); "Armeniâ Rossicá (Buhse!)" (cf. Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Russian Armenia" (cf. Stapf, 1915: 288, sub Phelypaea orientalis, flore coccinea); "Calculated" (cf. Domina & Raab-Straube, 2010, sub Ph. tournefortii).

Khojavend / Xocavənd [unconfirmed]: "Tale Dschagritschai, bei Takjalta Kulu [Dağdöşü / Tak] (Bayern!)" (cf. Beck, 1930: 44, sub Ph. tournefortii).

Nakhchivan Autonomous Republic / Naxçıvan Muxtar Respublikası: "444. Anoplanthus Tournefortii / Walp. ex parte / Puskoh bei Nachitschewan / 31/V, 47 [31 May 1847] / Buhse [leg.] // Iter Persicum. / Buhse, 1847-1849 / Herb. Al. de Bunge" (P 02970936 - E. Cosson, rev. Sánchez Pedraja, 30 Nov. 2011 [the two specimens on the upper left corner close to Buhse's label]); "Transcaucasia: Near Pusko in the Dshagri valley, north of Nahitshewan [Nachitschewan / Nakhchivan], on Pyrethrum myriophyllum, C.A. Mey., Buhse" (cf. Stapf, 1915: 293, sub P. tournefortii); "in der Provinz Nachitschevan [Nakhchivan Autonomous Republic] um Nachitschevan [Nakhchivan / Naxçıvan] bei Puskoh (Buhse!)" (cf. Beck, 1930: 44, sub Ph. tournefortii).

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Georgia / Georgien

"in pratis montosis Georgiae Caucasicae (Hoh!)" (cf. Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis [see Georgia in Reuter, 1847: 42, sub Anoplanthus biebersteinii Reut.]); "Georgien (Hohenacker nach Boissier)" (cf. Beck, 1930: 44); Georgian Biodiversity Database, accessed 11 Dec. 2010).

"Anoplon Biebersteinii C.A.M. / Caucas / 1850. / leg. Kovalevsky" (P 02970929, rev. Sánchez Pedraja, 30 Nov. 2011 [only one specimen well preserved, "calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes; scale-leaves concentrated towards the bases of the stems" (cf. Cullen, 2010: 384)]).

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Turkey / Türkiye / Tuquía / Turkije

Eastern Anatolia Region (Doğu Anadolu Bölgesi)

Ağrı Province: "B9 Ağri: Murat valley, 1600 m, D. 44038!" (cf. Gilli, 1982: 2, sub Ph. tournefortii).

Erzurum Province: Armenia (cf. Desfontaines, 1808: 18, sub Phelypaea orientalis, flore coccinea; Stapf, 1915: 285, 288, sub Phelypaea orientalis, flore coccinea, "native of Armenia"); "H. Native of Armenia" (cf. Don, 1838: 633, sub Anoplon tournefortii); "In Armeniâ (Tournef.)" (cf. Reuter, 1847: 43, sub Anoplanthus tournefortii); "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853 Armenia" (P 02970936 - E. Cosson, rev. Sánchez Pedraja, 30 Nov. 2011 [only one specimen on the lover left corner close to Huet de Pavillon's label]); A) ibidem, B) "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (P 02970941, rev. Sánchez Pedraja, 30 Nov. 2011 [six specimens very well preserved]); "Anoplon Tournefortii / ou / Phelipaea coccinea // 55. Top Dagh pres [?] / Erzeroum [Erzurum], Arménie. / Simon, 1854" (P 02970940 - E. Cosson, rev. Sánchez Pedraja, 30 Nov. 2011 [seven specimes well preserved]); A) "Orobanchea / Anoplon Tournefortii / Syn.:? / Phelipaea coccinea?", B) "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par M^r. le C^te. de Challaye 1855" (P 02970943, rev. Sánchez Pedraja, 30 Nov. 2011); "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, terra class. of Ph. tournefortii Desf.] (Tourn!, Huet!, Calv!)" (cf. Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "In Kurdistaniâ [Turkish Kurdistan] (J. Brant! Strangways in h. Benth.!)" (cf. Reuter, 1847: 43, sub Anoplanthus tournefortii); "Kurdistaniâ (Barré de Lancy [vice-consul de France à Mersine et Adana])!" (cf. Boissier, 1879: 494, sub Anoplanthus coccineus var. peduncularis); "discovered in Armenia, probably in the neighbourhood of Erzerum by Tournefort in 1701 [June 15 to July 6]" (cf. Stapf, 1915: 285, 289, 293, sub Phelypaea orientalis, flore coccinea and sub P. tournefortii); "Turkish Kurdistan" (cf. Stapf, 1915: 288, sub Phelypaea orientalis, flore coccinea); "Turkish Armenia: Upper Chorok valley, near Masut [Çakılı], E. of Baibut [Baiburt / Bayburt], 1200-1800 m, Huet de Pavillon! Near Erzerum, Zohrab, 826! Calver and Zohrab, 158! Kurdistan; without precise locality, Brant! Barre de Lancy (Hb. Gaillardot, 2555)" (cf. Stapf, 1915: 293, sub P. tournefortii); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum (Tournefort, Calvert nach Boissier); im Tale Kussuklu zwischen Baiburt und Erzerum bei 4000-5000' (Huet de Pavillon!) ... Kurdistan (Barre de Lancy nach Boissier)" (cf. Beck, 1930: 44, sub Ph. tournefortii); "E. Anatolia. A8 Erzurum: Sari Koma pass, 2600 m, Furse & Synge 850! Gümüşane: Kussuklu valley, Bayburt to Erzurum, 1220-1525 m, Huet." (cf. Gilli, 1982: 2, sub Ph. tournefortii); "AY209285: Turkey, NE Anatolia, between Bayburt and Erzurum, between the village Maden and Kop Geçidi, 1840 m, N 40° 04' 49'' E 40° 25' 34'', on Tanacetum sp. (Asteraceae), G. Schneeweiss & C.-G. Jang (WU: S&T 7764) (cf. Carlón & al., 2005: 63 sub Ph. tournefortii Desf.)"; "Gümüşane, Mescit Daglari: an der Straße zwischen Bayburt und Erzurum, zwischen dem Ort Maden und dem (Pass) Kop Geçidi; 40°04'49'' N, 40°25'34'' E. 1840 m, Schneeweiß, G. & Jang, C.-G. 7764 - AY209285, 2002-06-15" (WU 32431); "Erzurum" (photo by Hayati Keser in www.treknature.com, accessed 4 March 2012).

Kars Province: "im Distr. Kaghyzman [Kagizman / Kağızman, Kars, Eastern Anatolia (Armenian Highlands)] am Sattel Kara dagh (S. Michailowsky)" (cf. Beck, 1930: 44, sub Ph. tournefortii); "A9 Kars: Yağmurlu Da., 2200 m, D. 30822!" (cf. Gilli, 1982: 2, sub Ph. tournefortii).

Van Province: "zwischen Van und Hokotz-vank (D. Tchitouny nach Aznavour 1917)" (cf. Beck, 1930: 44, sub Ph. tournefortii); "Van: between Van and Hokotz church, Tchitouny" (cf. Gilli, 1982: 2, sub Ph. tournefortii); "June 13, 2002. Near Caldiran [Van], eastern Turkey" (sub Ph. coccinea, 2006 Old Dominion University, accessed 11 Dec. 2010); Van: steppes le long du chemin d’accès aux chutes de Muradiye, 1826 m, UTM 38S 0392562 / 4324169, leg. P. Coulot & al., 24/05/2009 [habit, flower, calyx, scale-leaves].

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